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Cettiidae

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Cettiidae
Cetti's warbler, Cettia cetti
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Superfamily: Aegithaloidea
Family: Cettiidae
Alström, Ericson, Olsson, & Sundberg, 2006
Genera

Scotocerca
Abroscopus
Phyllergates
Tickellia
Horornis
Tesia
Cettia
Urosphena
Hemitesia

Cettiidae is a newly validated family of small insectivorous songbirds ("warblers"), formerly placed in the Old World warbler "wastebin" assemblage. It contains the typical bush warblers (Cettia) and their relatives. As a common name, cettiid warblers is usually used.[1]

Some taxonomic authorities include this entire family, along with the related genera Erythrocercus and Scotocerca, in an enlarged family Scotocercidae.[2]

Its members occur mainly in Asia, ranging into Oceania and Europe. The pseudo-tailorbirds, tesias and stubtails, as well as Tickellia and Abroscopus warblers, are mostly found in the forests of south and southeastern Asia, with one species of stubtail reaching as far north as Japan and Siberia. Only one species, Neumann's warbler (Hemitesia neumanni), occurs in Africa.[3] The range of the genus Cettia extends west to Western Europe, while that of Horornis extends well into the Pacific, as far as Fiji and Palau. Most of the species in the family are sedentary, but the Asian stubtail is wholly migratory and the Japanese bush warbler and Cetti's warbler are partly migratory over much of their range. A few species, such as the pale-footed bush warbler, are altitudinal migrants.

The species are small, stubby birds. Most have moderately long to long tails, while the stubtails and tesias have tiny tails that do not even emerge past their tail coverts. The group is typically clad in dull plumage, often with a line above the eye, but some are more colorful. Altogether the Cettiidae are a quite variable group containing many aberrant birds that hitherto had been uncomfortably placed with a wide range of unrelated families.[citation needed] The Locustellidae, which contain birds which appear very similar to many cettiids, are far more uniform by contrast.[4]

Most live in scrubland and frequently hunt food by clambering through thick tangled growth.

Taxonomy

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The family Cettiidae was introduced by Per Alström and coworkers in 2006.[1]

The following cladogram showing the family relationships is based on a study by Carl Oliveros and coworkers published in 2019.[5] The number of species is taken from the AviList checklist.[6]

Aegithaloidea

Phylloscopidae – leaf warblers (80 species)

Hyliidae – hylias (2 species)

Aegithalidae – bushtits (11 species)

Erythrocercidae – bristle flycatchers (3 species)

Cettiidae – bush warblers and allies (32 species)

The phylogenetic relationships between eight of the genera were determined in a 2011 study by Per Alström and coworkers.[7] The streaked scrub warbler was not sampled in this study. The basal position shown here is based on the 2019 study by Carl Oliveros and coworkers. The streaked scrub warbler is estimated to have shared a common ancestor with the rest of the family around 16 million years ago.[5]

Cettiidae

Scotocerca – streaked scrub warbler

Hemitesia – (2 species)

Urosphena – stubtails (3 species)

Tesia – tesias (4 species)

Cettia – (4 species)

Abroscopus – warblers (3 species)

Phyllergates – tailorbirds (2 species)

Tickellia – Broad-billed warbler

Horornis – bush warblers (13 species)

The family contains 32 species in nine genera.[6]

Image Genus Living species
Scotocerca (Cretzschmar, 1830)
Abroscopus Baker, 1930 – warblers
Phyllergates Sharpe, 1883 – tailorbirds
Tickellia Blyth, 1861 – broad-billed warbler
Horornis Hodgson, 1845 – bush warblers
Tesia Hodgson, 1837 – tesias
Cettia Bonaparte, 1834 – typical bush warblers
Urosphena R. Swinhoe, 1877 – stubtails
Hemitesia Chapin, 1948 – warblers

References

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  1. ^ a b Alström, P.; Ericson, P.G.P.; Olsson, U.; Sundberg, P. (2006). "Phylogeny and classification of the avian superfamily Sylvioidea". Molecular Phylogenetics and Evolution. 38 (2): 381–397. doi:10.1016/j.ympev.2005.05.015.
  2. ^ Winkler D.; Billerman, S.; Lovette I. (2015). Bird Families of the World. Lynx Editions. pp. 442–444. ISBN 978-84-941892-0-3.
  3. ^ Irestedt, M.; Gelang, M.; Sangster, G.; Olsson, U.; Ericson, P.G.P.; Alström, P. (2011). "Neumann's Warbler Hemitesia neumanni (Sylvioidea): the sole African member of a Palaeotropic Miocene avifauna". Ibis. 153 (1): 78–86. doi:10.1111/j.1474-919X.2010.01084.x.
  4. ^ del Hoyo et al. (2006)
  5. ^ a b Oliveros, C.H.; et al. (2019). "Earth history and the passerine superradiation". Proceedings of the National Academy of Sciences of the United States of America. 116 (16): 7916–7925. doi:10.1073/pnas.1813206116. PMC 6475423. PMID 30936315.
  6. ^ a b AviList Core Team (2025). "AviList: The Global Avian Checklist, v2025". doi:10.2173/avilist.v2025. Retrieved 23 November 2025.
  7. ^ Alström, P.; Höhna, S.; Gelang, M.; Ericson, P.G.; Olsson, U. (2011). "Non-monophyly and intricate morphological evolution within the avian family Cettiidae revealed by multilocus analysis of a taxonomically densely sampled dataset". BMC Evolutionary Biology. 11 (1): 352. doi:10.1186/1471-2148-11-352. PMC 3261208.

Sources

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  • del Hoyo, Josep; Elliott, Andrew & Sargatal, Jordi (eds.) (2006): Handbook of Birds of the World (Volume 11: Old World Flycatchers to Old World Warblers). Lynx Edicions, Barcelona. ISBN 84-96553-06-X
  • Sefc, K.M.; Payne, R.B. & Sorenson, M.D. (2003): Phylogenetic relationships of African sunbird-like warblers: Moho Hypergerus atriceps, Green Hylia Hylia prasina and Tit-hylia Pholidornis rushiae. Ostrich 74(1-2): 8-17. PDF fulltext