Harkless Formation
Stratigraphic range: Lower Cambrian [1]
TypeFormation
Sub-unitsSee: Members
Underlies
OverliesPoleta Formation
Thickness0–3,600 ft (0–1,097 m)[1]
Lithology
PrimarySiltstone
OtherQuartzite, Limestone
Location
RegionNevada and California
CountryUnited States
Type section
Named forHarkless Flat

The Harkless Formation is a geologic formation in Nevada and California. It preserves fossils dating back to the Cambrian period.[1][2]

It is named from exposures on a divide south of Harkless Flat in the southern half of the Waucoba Mountain 15-min quadrangle in the Inyo Mountains of central eastern California.[3]

Geology

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The Harkless Formation is mainly composed of siltstones, with many layers of fine to medium-grained quartzites, with limestone within the lower parts of the formation, although differs in various parts of the formation.[1] In the Split Mountain area, the formation underlies the Mule Spring Limestone,[1][4] whilst in other areas it underlies the Saline Valley Formation.[1]

Members

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The formation contains two members,[4] which are as follows, in ascending age:

  • Weepah Member: Near the Andrews Mountain, at the type area of the formation, the member is dominated by greenish-gray and light-olive-gray siltstone, alongside a few layers of yellowish-gray fine to medium grained quartzite. The quartzite layers are identical to the layers found in the Zabriskie Quartzite. The siltstone ranges from grayish-olive, pale-olive, and dark-greenish-gray, and is composed of silt-size grains within a matrix of muscovite and chlorite, and is thinly laminated. They have also slightly metamorphosed, resulting in phyllite or hornfels. The hornfels are primarily composed of muscovite, chlorite, biotite, and quartz. Trace fossils are relatively common within the type area.[1] Within the Esmeralda County, Nevada, the lower sections of the member is still dominated by siltstone, although this differs greatly from the siltstones higher up in the formation. This is due to the layers being quartzitic, coarser and more resistant to natural erosion. The sizes of the grains within these layers hovers around the boundary between coarse silt and very fine sand.[1]
  • Alkali Member: The upper Alkali member is composed of limestone layers, up to 100 ft (30 m) thick, inter-stratified with siltstone, and contain an abundance of archeocyathid sponges. Further still there are light-brown limestone layers, only up to 1 ft (0.30 m) thick, that contain Salterella, and are also inter-stratified with the siltstone layers in the middle of the formation. More limestone can be found at the top of the member, being gray or locally yellowish-brown, and up to 40 ft (12 m) thick.[1] In some areas of the member, there are also tongues from the Zabriskie Quartzite, composed primarily of medium-grained quartzite, and can get up to 300 ft (91 m) thick in some places, and inter-stratify with the siltstones. The colour of the quartzite is commonly yellowish-gray or greenish-gray. It is also laminated to thin-bedded. Trace fossils are also common within these quartzite tongues, also appearing on the underlying siltstone layers.[1]

Paleobiota

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The Harkless Formation contains a number of sponge spicules, hyoliths, brachiopods, and other small shelly fauna.[5][2] Alongside this, there are a myriad of arthropods, namely trilobites,[6] including a single bradoriid, Mongolitubulus.[7]

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Arthropoda

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Genus Species Notes Images
Mesonacis[6]
  • M. fremonti
Olenellid trilobite, originally described as Fremonita.[8]
Olenellus[1][6][4]
  • Olenellus sp.
  • O. clarki
  • O. glabrum
Olenellid trilobite.
Wanneria[9]
  • Wanneria sp.
Olenellid trilobite.
Ogygopsis[6][4]
  • Ogygopsis sp.
Dorypygid trilobite.
Bonnia[6][4]
  • Bonnia sp.
  • B. brennus
  • B. columbensis
Dorypygid trilobite.
Ovatoryctocara[4]
  • O. yaxiensis
Corynexochid trilobite.
Oryctocephalops[4]
  • O. frischenfeldi
Corynexochid trilobite.
Zacanthopsis[4]
  • Zacanthopsisi sp.
  • Z. levis
Corynexochid trilobite.
Protoryctocephalus (?)[4]
  • P. articus (?)
Corynexochid trilobite.
Bristolia[6][4]
  • B. bristolensis
  • B. mohavensis
Biceratopsid trilobite, previously reported as Olenellus gilberti.[1]
Anebocephalus[4]
  • A. silverpeakensis
Trilobite, family unknown.
Coenoides[4]
  • C. scholteni
Trilobite, family unknown.
Harklessaspis[4]
  • H. rasettii
  • H. parvigranulosus
Trilobite, family unknown.
Arcuolenellus[4]
  • A. megafrontalis
Trilobite, family unknown.
Crassifimbra[4]
  • Crassifimbra sp.
  • C. walcotti
Trilobite, family unknown.
Mongolitubulus[7]
  • M. squamifer
Bradoriid arthropod.

Lophotrochozoa

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Genus Species Notes Images
Hyolithellus[5][2]
  • H. insolitus
Lophotrochozoan hyolith. May possibly be an annelid.[10]
Kyrshabaktella[5][7]
  • K. hicksi
Linguliform brachiopod.
Eothele (?)[5]
  • E. spurri (?)
Acrotheloid brachiopod.
Hadrotreta[5][7]
  • H. primaea
Acrotheloid brachiopod.
Obolella[11]
  • Obolella sp.
Obelellid brachiopod.
Mickwitzia[11]
  • M. occidens
Mickwitziid brachiopod.
Micromitra[7]
  • Micromitra sp.
Paterinid brachiopod.
Anuliconus[7]
  • Anuliconus sp.
Helcionellid mollusc.
Davidonia (?)[7]
  • D. rostrata (?)
Helcionellid mollusc.
Pelagiella[9][7]
  • P. subangulata
Pelagiellid mollusc.
Harkless bryomorph[12]
  • ???
Probable bryozoan, though this has been contested.[13]

Chancelloriidae

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Genus Species Notes Images
Allonia[7]
  • Allonia sp.
Chancelloriid.
Chancelloria[7]
  • Chancelloria sp.
Chancelloriid.

Cnidaria

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Genus Species Notes Images
Sphenothallus[5][2]
  • Sphenothallus sp.
Cnidarian conulariid.

Echinodermata

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Genus Species Notes Images
Echinoderm Ossicles[5][2][7]
  • ???
Calcareous element that makes up the endoskeleton of echinoderms.

Porifera (Sponges)

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Genus Species Notes Images
Retilamina[9]
  • R. debrenneae
Archeocyathid sponge.
Robertiolynthus[9]
  • R. handfieldi
Archeocyathid sponge.
Cjulanciella (?)[7]
  • C. asimmetrica (?)
Hexactinellid sponges.
Archeocyathids[1][7]
  • ???
Archeocyathid sponge fragments, found in the lower layers of the formation.
Hexactinellid spicules[5][9]
  • ???
Sponge skeleton, with four rays.
Heteractinida spicules[5][9][7]
  • ???
Sponge skeleton, composed of four rays radiating outwards, and two rays rising up from the plane.

incertae sedis

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Genus Species Notes Images
Volborthella[6]
  • Volborthella sp.
Tubular salterellid fossil.
Salterella[1][6][7]
  • S. conulata
Tubular salterellid fossil.
Lidaconus[14]
  • L. palmettoensis
Tubular fossil.

Flora

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Genus Species Notes Images
Renalcid-like[9]
  • ???
Cyanophyte cyanobacteria.

Ichnogenera

[edit]
Genus Species Notes Images
Cruziana[6]
  • Cruziana sp.
Trails.
Planolites[6]
  • Planolites sp.
Trails.
Scolicia[6]
  • Scolicia sp.
Burrows.
Monocraterion[6]
  • Monocraterion sp.
Burrows.
Rusophycus[6]
  • Rusophycus sp.
  • R. didymus
Resting trace of trilobites.
Skolithos[6]
  • Skolithos sp.
Burrows.
Bergaueria[6]
  • Bergaueria sp.
Resting trace of cnidarians.
Diplichnites[6]
  • Diplichnites sp.
Arthropod trackways.
Archaeonassa[6]
  • Archaeonassa sp.
Burrows.
Monomorphichnus[6]
  • M. multilineatus
Trilobite feeding traces.
Asteriacites (?)[6]
  • Asteriacites (?) sp.
Sea star burrows.

See also

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References

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  1. ^ a b c d e f g h i j k l m n Stewart, J. H. "Upper Precambrian and Lower Cambrian Strata, in the Southern Great Basin California and Nevada" (PDF). U.S. Geological Survery. Geological Survey Professional.
  2. ^ a b c d e Wotte, Thomas; Sundberg, Frederick A. (September 2017). "Small shelly fossils from the Montezuman–Delamaran of the Great Basin in Nevada and California". Journal of Paleontology. 91 (5): 883–901. doi:10.1017/jpa.2017.8.
  3. ^ "Geologic Unit: Harkless". National Geologic Map Database. U.S. Geological Survey. Retrieved 23 April 2022.
  4. ^ a b c d e f g h i j k l m n o Sundberg, Frederick A.; Webster, Mark (July 2022). ""Ptychoparioid" trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada". Journal of Paleontology. 96 (4): 886–920. doi:10.1017/jpa.2021.124.
  5. ^ a b c d e f g h i Skovsted, Christian B.; Holmer, Lars E. (December 2006). "The Lower Cambrian brachiopod Kyrshabaktella and associated shelly fossils from the Harkless Formation, southern Nevada". GFF. 128 (4): 327–337. doi:10.1080/11035890601284327.
  6. ^ a b c d e f g h i j k l m n o p q r s Alpert, Stephen P. (1976). "Trilobite and Star-like Trace Fossils from the White-Inyo Mountains, California". Journal of Paleontology. 50 (2): 226–239. ISSN 0022-3360.
  7. ^ a b c d e f g h i j k l m n o Bennett, Casey C.; Mate, Clare; Jacquet, Sarah M. (July 2025). "Small shelly fauna biodiversity from reef-adjacent facies of the lower Cambrian Harkless Formation, Nevada". Palaeogeography, Palaeoclimatology, Palaeoecology. 669 112922. doi:10.1016/j.palaeo.2025.112922.
  8. ^ Firby, Jean B.; Durham, J. Wyatt (1974). "Molluscan Radula from Earliest Cambrian". Journal of Paleontology. 48 (6): 1109–1119. ISSN 0022-3360.
  9. ^ a b c d e f g Pruss, Sara B.; Smith, Emily F.; Leadbetter, Olivia; Nolan, Rhiannon Z.; Hicks, Melissa; Fike, David A. (December 2019). "Palaeoecology of the archaeocyathan reefs from the lower Cambrian Harkless Formation, southern Nevada, western United States and carbon isotopic evidence for their demise". Palaeogeography, Palaeoclimatology, Palaeoecology. 536 109389. doi:10.1016/j.palaeo.2019.109389.
  10. ^ Skovsted, Christian B.; Peel, John S. (January 2011). "Hyolithellus in life position from the Lower Cambrian of North Greenland". Journal of Paleontology. 85 (1): 37–47. doi:10.1666/10-065.1.
  11. ^ a b Rowell, A. J. (1977). "Early Cambrian Brachiopods from the Southwestern Great Basin of California and Nevada". Journal of Paleontology. 51 (1): 68–85. ISSN 0022-3360.
  12. ^ Pruss, Sara B.; Leeser, Lexie; Smith, Emily F.; Zhuravlev, Andrey Yu.; Taylor, Paul D. (22 April 2022). "The oldest mineralized bryozoan? A possible palaeostomate in the lower Cambrian of Nevada, USA". Science Advances. 8 (16). doi:10.1126/sciadv.abm8465. PMC 9020656.
  13. ^ Yang, Jie; Lan, Tian; Zhang, Xi-guang; Smith, Martin R. (16 March 2023). "Protomelission is an early dasyclad alga and not a Cambrian bryozoan". Nature. 615 (7952): 468–471. doi:10.1038/s41586-023-05775-5.
  14. ^ Onken, Beth R.; Signor, Philip W. (1988). "Lidaconus palmettoensis n. gen. and sp.: An Enigmatic Early Cambrian Fossil from Western Nevada". Journal of Paleontology. 62 (2): 172–180. ISSN 0022-3360.