↑Jordan, D. S. & H. W. Fowler, 1903. A review of the cyprinoid fishes of Japan. Proceedings of the United States National Museum v. 26 (núm. 1334): 811-862.
↑Oshima, M., 1957. Notes on the fishes collected in the down stream of Tama River, with descriptions of two new species. Japanese Journal of Ichthyology v. 6 (núms. 1-2): 14-19.