(en) Bastian Bentlage, Paulyn Cartwright, Angel A Yanagihara, Cheryl Lewis, Gemma S. Richards et Allen G Collins, « Evolution of box jellyfish (Cnidaria: Cubozoa), a group of highly toxic invertebrates », Proceedings of the Royal Society B, Royal Society, vol. 277, no 1680, , p. 493-501 (ISSN0962-8452 et 1471-2954, PMID19923131, PMCID2842657, DOI10.1098/RSPB.2009.1707, lire en ligne).