(en) V. Dincă, V. A. Lukhtanov, G. Talavera et R. Vila, « Unexpected layers of cryptic diversity in wood white Leptidea butterflies », Nature communications, vol. 2, no 324, (DOI10.1038/ncomms1329).
(en) V. Dincă, C. Wiklund, V. A. Lukhtanov, U. Kodandaramaiah, K. Norén, L. Dapporto, N. Wahlberg, R. Vila et M. Friberg, « Reproductive isolation and patterns of genetic differentiation in a cryptic butterfly species complex », Journal of Evolutionary Biology, vol. 26, no 10, , p. 2095–2106 (DOI10.1111/jeb.12211).