↑ abc et dChang, C.-H., Li, F., Shao, K.-T., Lin, Y.-S., Morosawa, T., Kim, S., Koo, H., Kim, W., Lee, J.-S., He, S., Smith, C., Reichard, M., Miya, M., Sado, T., Uehara, K., Lavoué, S., Chen, W.-J. & Mayden, R.L., « Phylogenetic relationships of Acheilognathidae (Cypriniformes: Cyprinoidea) as revealed from evidence of both nuclear and mitochondrial gene sequence variation: Evidence for necessary taxonomic revision in the family and the identification of cryptic species », Molecular Phylogenetics and Evolution,, vol. 81, , p. 182–194. (DOI10.1016/j.ympev.2014.08.026, lire en ligne)