ハプティスタ (学名 :Haptista)は、ハプト藻 や有中心粒太陽虫 から構成される原生生物 のグループとして提案されている。系統発生学の研究では、ハプティスタはSARスーパーグループ の姉妹群 を形成していることを示している[ 1] [ 2] 。
2015年 のキャバリエ=スミス 及びChao & Lewisの研究[ 3] と2015年のRuggiero et al. の研究[ 4] に基づいた分類を下記に示す。
亜門 Centroheliozoa Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987 [Heliozoa Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003 ]
綱 Centrohelea Kuhn 1926 stat. n. Cavalier-Smith 1993 [Centroplastiales; Centrohelina Hartmann 1913 ; Centroplasthelida Febvre-Chevalier 1984 ]
亜門 ハプト藻 Haptophytina Cavalier-Smith 2015 (Haptophyta Hibberd 1976 sensu Ruggerio et al. 2015 )
クレード Rappemonada Kim et al. 2011
クレード Haptomonada (Margulis & Schwartz 1998) [Haptophyta Hibberd 1976 emend. Edvardsen & Eikrem 2000; Prymnesiophyta Green & Jordan, 1994 ; Prymnesiomonada; Prymnesiida Hibberd 1976; Haptophyceae Christensen 1962 ex Silva 1980; Haptomonadida; Patelliferea Cavalier-Smith 1993]
綱 Pavlovophyceae (Cavalier-Smith 1986) Green & Medlin 2000
綱 Prymnesiophyceae Christensen 1962 emend. Cavalier-Smith 1996 [Haptophyceae s.s.; Prymnesiophycidae Cavalier-Smith 1986 ; Coccolithophyceae Casper 1972 ex Rothmaler 1951 ]
亜門 Alveidia Cavalier-Smith 2018 [ 5]
^ Burki F; Kaplan M; Tikhonenkov DV; Zlatogursky V; Minh BQ; Radaykina LV; Smirnov A; Mylnikov AP et al. (2016), “Untangling the early diversification of eukaryotes: a phylogenomic study of the evolutionary origins of Centrohelida, Haptophyta and Cryptista.” , Proc Biol Sci 283 (1823): 20152802, doi :10.1098/rspb.2015.2802 , PMC 4795036 , PMID 26817772 , http://www.pubmedcentral.nih.gov/articlerender.fcgi?tool=pmcentrez&artid=4795036
^ Burki, F; Inagaki, Y; Bråte, J; Archibald, J.; Keeling, P.; Cavalier-Smith, T; Sakaguchi, M; Hashimoto, T et al. (2009). “Large-scale phylogenomic analyses reveal that two enigmatic protist lineages, Telonemia and Centroheliozoa, are related to photosynthetic chromalveolates.” (Free full text). Genome Biology and Evolution 1 : 231–8. doi :10.1093/gbe/evp022 . PMC 2817417 . PMID 20333193 . オリジナル の2012-07-10時点におけるアーカイブ。. https://archive.is/20120710011541/http://gbe.oxfordjournals.org/cgi/content/short/evp022v1?rss=1 .
^ Cavalier-Smith; Chao; Lewis (2015), “Multiple origins of Heliozoa from flagellate ancestors: New cryptist subphylum Corbihelia, superclass Corbistoma, and monophyly of Haptista, Cryptista, Hacrobia and Chromista”, Molecular Phylogenetics and Evolution 93 : 331–362, doi :10.1016/j.ympev.2015.07.004 , PMID 26234272
^ Ruggiero (2015), “Higher Level Classification of All Living Organisms” , PLoS ONE 10 (4): e0119248, Bibcode : 2015PLoSO..1019248R , doi :10.1371/journal.pone.0119248 , PMC 4418965 , PMID 25923521 , http://www.pubmedcentral.nih.gov/articlerender.fcgi?tool=pmcentrez&artid=4418965
^
Cavalier-Smith, T.; Chao, E. E.; Lewis, R. (17 April 2018), “Multigene phylogeny and cell evolution of chromist infrakingdom Rhizaria: contrasting cell organisation of sister phyla Cercozoa and Retaria” , Protoplasma 255 (5): 1517–1574, doi :10.1007/s00709-018-1241-1 , PMC 6133090 , PMID 29666938 , http://www.pubmedcentral.nih.gov/articlerender.fcgi?tool=pmcentrez&artid=6133090