꼭두서니아과

꼭두서니아과
Rubia tinctorum
생물 분류ℹ️
계:	식물계
(미분류):	속씨식물군
(미분류):	진정쌍떡잎식물군
(미분류):	국화군
목:	용담목
과:	꼭두서니과
아과:	꼭두서니아과 Rubioideae Verdc., 1958
모식종
꼭두서니속(Rubia L.)
^[출처 필요]
속
본문 참조

꼭두서니아과(----亞科, 학명: Rubioideae 루비오이데아이^[*])는 꼭두서니과의 아과이다.^[1] 26개 족, 약 7600여 종으로 이루어져 있다.^[2][3] 갈퀴덩굴, 계요등, 꼭두서니, 노니, 민둥갈퀴, 솔나물, 수정목, 호자나무 등을 포함하고 있다.

하위 분류

가이르트네라족(Gaertnereae Endl.)^[4]

• Gaertnera Lam.
• Pagamea Aubl.

계요등족(Paederieae DC.)^[5]

• 계요등속(Paederia L.)
• Leptodermis Wall.
• Pseudopyxis Miq.
• Saprosma Blume
• Serissa Comm. ex A.Juss.
• Spermadictyon Roxb.

꼭두서니족(Rubieae Baill.)^[6]

• 갈퀴덩굴속(Galium L.)
• 갈퀴아재비속(Asperula L.)
• 꼭두서니속(Rubia L.)
• 꽃갈퀴덩굴속(Sherardia L.)
• Callipeltis Steven
• Crucianella L.
• Cruciata Mill.
• Didymaea Hook.f.
• × Galiasperula Ronniger
• Kelloggia Torr. ex Benth. & Hook.f.
• Mericarpaea Boiss.
• Microphysa Schrenk
• Phuopsis (Griseb.) Hook.f.
• Valantia L.

노니족(Morindeae Kostel.)^[7]

• 노니속(Morinda L.)
• Appunia Hook.f.
• Coelopyrena Valeton
• Coelospermum Blume
• Gynochthodes Blume

다나이스족(Danaideae B.Bremer & Manen)^[8]

• Danais Comm. ex Vent.
• Payera Baill.
• Schismatoclada Baker

둔니아족(Dunnieae Rydin & B.Bremer)^[9]

• Dunnia Tutcher

무주나무족(Lasiantheae B.Bremer & Manen)^[10]

• 무주나무속(Lasianthus Jack)
• Ronabea Aubl.
• Saldinia A.Rich. ex DC.
• Trichostachys Hook.f.

스크라데라족(Schradereae Bremek.)^[11]

• Lecananthus Jack
• Leucocodon Gardner
• Schradera Vahl

스키조콜레아족(Schizocoleeae Rydin & B.Bremer)^[12]

• Schizocolea Bremek.

스페르마코케족(Spermacoceae Bercht. & J.Presl)^[13]

• 낚시돌풀속(Leptopetalum Hook. & Arn.)
• 백령풀속(Hexasepalum Bartl. ex DC.)
• 백운풀속(Scleromitrion (Wight & Arn.) Meisn.)
• 제주백운풀속(Oldenlandia L.)
• 큰백령풀속(Diodia L.)
• 탐라풀속(Neanotis W.H.Lewis)
• Agathisanthemum Klotzsch
• Amphiasma Bremek.
• Amphistemon Groeninckx
• Anthospermopsis (K.Schum.) J.H.Kirkbr.
• Arcytophyllum Willd.
• Astiella Jovet
• Bouvardia Salisb.
• Carajasia R.M.Salas, E.L.Cabral & Dessein
• Carterella Terrell
• Conostomium (Stapf) Cufod.
• Cordylostigma Groeninckx & Dessein
• Crusea Schltdl. & Cham.
• Debia Neupane & N.Wikstr.
• Denscantia E.L.Cabral & Bacigalupo
• Dentella J.R.Forst. & G.Forst.
• Diacrodon Sprague
• Dibrachionostylus Bremek.
• Dimetia (Wight & Arn.) Meisn.
• Edrastima Raf.
• Eleuthranthes F.Muell. ex Benth.
• Emmeorhiza Pohl ex Endl.
• Ernodea Sw.
• Exallage Bremek.
• Galianthe Griseb.
• Gomphocalyx Baker
• Hedyotis L.
• Hedythyrsus Bremek.
• Houstonia L.
• Hydrophylax L.f.
• Involucrella (Benth. & Hook.f.) Neupane & N.Wikstr.
• Kadua Cham. & Schltdl.
• Kohautia Cham. & Schltdl.
• Lelya Bremek.
• Leptomischus Drake
• Manettia Mutis ex L.
• Manostachya Bremek.
• Mexotis Terrell & H.Rob.
• Mitracarpus Zucc.
• Mitrasacmopsis Jovet
• Nesohedyotis (Hook.f.) Bremek.
• Oldenlandiopsis Terrell & W.H.Lewis
• Opercularia Gaertn.
• Pentanopsis Rendle
• Pentodon Hochst.
• Phialiphora Groeninckx
• Phyllocrater Wernham
• Phylohydrax Puff
• Planaltina R.M.Salas & E.L.Cabral
• Pseudonesohedyotis Tennant
• Psyllocarpus Mart. & Zucc.
• Richardia L.
• Sacosperma G.Taylor
• Schwendenera K.Schum.
• Spermacoce L.
• Staelia Cham. & Schltdl.
• Stenaria (Raf.) Terrell
• Stenotis Terrell
• Synaptantha Hook.f.
• Tessiera DC.
• Thamnoldenlandia Groeninckx

아르고스템마족(Argostemmateae Bremek. ex Verdc.)^[14]

• Argostemma Wall.
• Mouretia Pit.
• Mycetia Reinw.
• Neohymenopogon Bennet

안토스페르뭄족(Anthospermeae Cham. & Schltdl.)^[15]

• Anthospermum L.
• Carpacoce Sond.
• Coprosma J.R.Forst. & G.Forst.
• Durringtonia R.J.F.Hend. & Guymer
• Galopina Thunb.
• Leptostigma Arn.
• Nenax Gaertn.
• Nertera Banks ex Gaertn.
• Normandia Hook.f.
• Phyllis L.
• Pomax Sol. ex DC.

오피오리자족(Ophiorrhizeae Bremek. ex Verdc.)^[16]

• Coptophyllum Korth.
• Keenania Hook.f.
• Lerchea L.
• Neurocalyx Hook.
• Ophiorrhiza L.
• Spiradiclis Blume
• Xanthophytum Reinw. ex Blume

우로필룸족(Urophylleae Bremek. ex Verdc.)^[17]

• Amphidasya Standl.
• Antherostele Bremek.
• Pauridiantha Hook.f.
• Pentaloncha Hook.f.
• Raritebe Wernham
• Temnopteryx Hook.f.
• Urophyllum Jack ex Wall.

코우사레아족(Coussareeae Hook.f.)^[18]

• Bradea Standl.
• Coccocypselum P.Browne
• Coussarea Aubl.
• Cruckshanksia Hook. & Arn.
• Declieuxia Kunth
• Faramea Aubl.
• Heterophyllaea Hook.f.
• Hindsia Benth. ex Lindl.
• Oreopolus Schltdl.

콜레토이케마족(Colletoecemateae Rydin & B.Bremer)^[19]

• Colletoecema E.M.A.Petit

크녹시아족(Knoxieae Benth. & Hook.f.)^[20]

• Batopedina Verdc.
• Carphalea Juss.
• Chamaepentas Bremek.
• Dirichletia Klotzsch
• Dolichopentas Kårehed & B.Bremer
• Knoxia L.
• Lathraeocarpa Bremek.
• Otiophora Zucc.
• Otomeria Benth.
• Paracarphalea Razafim., Ferm, B.Bremer & Kårehed
• Paraknoxia Bremek.
• Parapentas Bremek.
• Pentanisia Harv.
• Pentas Benth.
• Phyllopentas (Verdc.) Kårehed & B.Bremer
• Rhodopentas Kårehed & B.Bremer
• Triainolepis Hook.f.

크라테리스페르뭄족(Craterispermeae Verdc.)^[21]

• Craterispermum Benth.

키아노네우론족(Cyanoneuroneae Razafim. & B.Bremer)^[22]

• Cyanoneuron Tange

텔리고눔족(Theligoneae Baill.)^[23]

• Theligonum L.

팔리코우레아족(Palicoureeae Robbr. & Manen)^[24]

• Carapichea Aubl.
• Chassalia Comm. ex Poir.
• Ditrichanthus Borhidi, E.Martínez & Ramos
• Eumachia DC.
• Geophila D.Don
• Hymenocoleus Robbr.
• Notopleura (Hook.f.) Bremek.
• Palicourea Aubl.
• Puffia Razafim. & B.Bremer
• Rudgea Salisb.
• Tromlyca Borhidi

페라마족(Perameae Bremek. ex S.P.Darwin)^[25]

• Perama Aubl.

퐁케비아족(Foonchewieae R.J.Wang)^[26]

• Foonchewia R.J.Wang

푸토리아족(Putorieae Sweet)^[27]

• Plocama Aiton

프리스마토메리스족(Prismatomerideae Y.Z.Ruan)^[28]

• Gentingia J.T.Johanss. & K.M.Wong
• Motleyia J.T.Johanss.
• Prismatomeris Thwaites
• Rennellia Korth.

프시코트리아족(Psychotrieae Cham. & Schltdl.)^[29]

• Amaracarpus Blume
• Anthorrhiza C.R.Huxley & Jebb
• Calycosia A.Gray
• Dolianthus C.H.Wright
• Hedstromia A.C.Sm.
• Hydnophytum Jack
• Myrmecodia Jack
• Myrmephytum Becc.
• Psychotria L.
• Squamellaria Becc.
• Streblosa Korth.

호자덩굴족(Mitchelleae Razafim. & B.Bremer)^[30]

• 호자나무속(Damnacanthus C.F.Gaertn.)
• 호자덩굴속(Mitchella L.)

분류 불확실(incertae sedis)

• Aphanocarpus Steyerm.
• Clarkella Hook.f.
• Coryphothamnus Steyerm.
• Pagameopsis Steyerm.

계통 분류

꼭두서니과[31]	꼭두서니아과       치자나무아과     구슬꽃나무아과         콥토사펠타족     루쿨리아족

각주

1. Verdcourt, Bernard. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 280. 1958.
2. Juss., 1789. Genera Plantarum 196.
3. B. Bremer and J.-F. Manen, Phylogeny and classification of the subfamily Rubioideae (Rubiaceae) Plant Syst. Evol. 225:43-72 (2000) PDF.
4. Endlicher, Stephan Friedrich Ladislaus. Genera Plantarum (Endlicher) 1: 577. 1838.
5. Candolle, Augustin Pyramus de. Prodromus Systematis Naturalis Regni Vegetabilis 4: 342, 470. 1830.
6. Baillon, Henri Ernest. Histoire des Plantes 7: 365, 390. 1880
7. Kosteletzky, Vincenz Franz. Allgemeine Medizinisch-Pharmazeutische Flora 2: 565. 1833.
8. Bremer, Birgitta & Manen, Jean-François. Plant Systematics and Evolution 225: 60. 2000.
9. Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 121. 2009.
10. Bremer, Birgitta & Manen, Jean-François. Plant Systematics and Evolution 225: 57. 2000.
11. Bremekamp, Cornelis Eliza Bertus. Recueil des Travaux Botaniques Néerlandais 31: 253. 1934.
12. Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 120. 2009.
13. Berchtold, Bedřich Všemír von & Presl, Jan Svatopluk. O Prirozenosti Rostlin 256. 1820.
14. Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.
15. Chamisso, Ludolf Karl Adelbert von & Schlechtendal, Diederich Franz Leonhard von. Linnaea 3: 309. 1828.
16. Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.
17. Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.
18. Hooker, Joseph Dalton. Genera Plantarum 2: 9. 1873.
19. Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 120. 2009.
20. Bentham, George & Hooker, Joseph Dalton. Genera Plantarum 2: 9, 21. 1873.
21. Verdcourt, Bernard. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.
22. Razafimandimbison, Sylvain Georges & Bremer, Birgitta. Taxon 64(2): 294. 2015.
23. Baillon, Henri Ernest. Histoire des Plantes 4: 46, 55. 1872.
24. Robbrecht, Elmar & Manen, Jean-François. Systematics and Geography of Plants 76: 125, 136. 2006.
25. Bremekamp, Cornelis Eliza Bertus. Taxon 25(5-6): 605. 1976.
26. Wang, Rui-jiang. Journal of Systematics and Evolution 50(5): 472–473. 2012.
27. Sweet, Robert. Hortus Britannicus, ed. 3 325. 1839.
28. Ruan, Yun Zhen. Acta Phytotaxonomica Sinica 26(6): 446. 1988.
29. Chamisso, Ludolf Karl Adelbert von & Schlechtendal, Diederich Franz Leonhard von. Linnaea 4(1): 4. 1829.
30. Razafimandimbison, Sylvain Georges & Bremer, Birgitta. Molecular Phylogenetics and Evolution 48(1): 221. 2008.
31. Bremer, Birgitta; Eriksson, Torsten (2009). “Time Tree of Rubiaceae: Phylogeny and Dating the Family, Subfamilies, and Tribes” (PDF). 《International Journal of Plant Sciences》 (영어) 170 (6): 766–793. doi:10.1086/599077.