↑Bell, N. E.; Quandt, D.; O'Brien, T. J.; Newton, A. E. 2007: Taxonomy and phylogeny in the earliest diverging pleurocarps: square holes and bifurcating pegs. Bryologist, 110: 533-560.
↑Goffinet, B.; Buck, W. R.; & Shaw, J. (2008) "Morphology and Classification of the Bryophyta". pp. 55-138 in Goffinet, B. & J. Shaw (eds.) Bryophyte Biology, 2nd ed. Cambridge University Press.