↑Reynolds,R.P. & Foster,M.S. (1992) Four new species of frogs and one new species of snake from the Chapare region of Bolivia, with notes on other species., Herpetological Monographs 6: 83-104
↑ 3.03.13.2Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)