↑ 2.02.12.2Randall, J.E. and P.C. Heemstra (1991) Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species., Indo-Pacific Fishes (20):332 p.
↑ 3.03.1Eschmeyer, W.N. (ed.) (2001) Catalog of fishes. Updated database version of December 2001., Catalog databases as made available to FishBase in December 2001.
↑Heemstra, P.C. (1995) Additions and corrections for the 1995 impression., p. v-xv. In M.M. Smith and P.C. Heemstra (eds.) Revised Edition of Smiths' Sea Fishes. Springer-Verlag, Berlin.
↑ 5.05.15.2Heemstra, P.C. and J.E. Randall (1993) FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date., FAO Fish. Synop. 125(16):382 p.
↑ 6.06.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011