↑ 2.02.1Nagelkerke, L.A.J. and F.A. Sibbing (1997) A revision of the large barbs (Barbus spp., Cyprinidae, Teleostei) of Lake Tana, Ethiopia, with a description of seven new species., p. 105-170. In L.A.J. Nagelkerke (ed.) The barbs of Lake Tana, Ethiopia: morphological diversity and its implications for taxonomy, trophic resource partititioning and fisheries. Wageningen University, The Netherlands.
↑ 3.03.1de Graaf, M., E.D. Nentwich, J.W.M. Osse and F.A. Sibbing (2005) Lacustrine spawning: is this a new reproductive strategy among 'large' African cyprinid fishes?., J. Fish Biol. 66(5):1214-1236.
↑ 4.04.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑FishBase. Froese R. & Pauly D. (eds), 14 Hunyo 2011