↑ 2.02.12.2Frost, Darrel R. (2009) , database. Amphibian Species of the World: an Online Reference v5.3
↑ 3.03.1(2004) , database, Amphibian Species of the World: an Online Reference
↑ 4.04.1Frost, Darrel R. (1997) , Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
↑Lynch, John D., and Juan M Renjifo (1990) Two New Toads (Bufonidae: Rhamphophryne) from the Northern Andes of Columbia, Journal of Herpetology, vol. 24, no. 4