Price, B.W., Villet, M.H., Walton, S.M., Barker, N.P. 2011. Using molecules and morphology to infer the phylogenetic relationships and evolutionary history of the Dirini (Nymphalidae: Satyrinae), a tribe of butterflies endemic to southern Africa. Systematic Entomology 36: 300-316.
寿建新 周尧 李宇飞. 世界蝴蝶分類名錄. 中國: 陝西科學技術出版社. 2006-04-01. ISBN 9787536936768(中文(简体)).