Beltrán, M., Jiggins, C. D., Brower, A. V. Z., Bermingham, E. & Mallet, J. 2007 Do pollen feeding, pupal-mating and larval gregariousness have a single origin in Heliconius butterflies? Inferences from multilocus DNA sequence data. Biol. J. Linn. Soc. 92, 221-239.
Brower, A. V. Z. & Egan, M. G. 1997 Cladistics of Heliconius butterflies and relatives (Nymphalidae: Heliconiiti): the phylogenetic position of Eueides based on sequences from mtDNA and a nuclear gene. Proc. R. Soc. Lond. B 264, 969-977.
Brown K. S. 1981 The Biology of Heliconius and Related Genera. Annual Review of Entomology 26, 427-456.
Creisler B. 1996 The Etymology of Chelodina Fitzinger, 1826 (Testudines: Chelidae. Journal of Herpetology, Vol. 30, No. 4 (Dec., 1996), pp. 556–556.
DeVries P. J. 1997 The Butterflies of Costa Rica and Their Natural History, Volume I: Papilionidae, Pieridae, Nymphalidae Princeton University Press, Baskerville, USA.