^ 1.01.1Wahlberg, N. 2006. That awkward age for butterflies: insights from the age of the butterfly subfamily Nymphalinae. Systematic Biology 55:703-714. (英文)
Kons, H. L. Jr. 2000: Phylogenetic studies of the Melitaeini (Lepidoptera: Nymphalidae: Nymphalinae) and a revision of the genus Chlosyne Butler. - PhD thesis, University of Florida, Gainesville, 799 pp. Available from UMI.
Wahlberg N, Brower AVZ, and Nylin S. 2005. Phylogenetic relationships of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae) based on three gene sequences. Biol. J. Linn. Soc. 86: 227-251.
Wahlberg, N. & Freitas, A. V. 2007 Colonization of and radiation in South America by butterflies in the subtribe Phyciodina (Lepidoptera: Nymphalidae). Molecular Phylogenetics and Evolution 44, 1257-1272.
Zimmermann, M., N. Wahlberg, and H. Descimon. 2000. Phylogeny of Euphydryas checkerspot butterflies (Lepidoptera: Nymphalidae) based on mitochondrial DNA sequence data. Annals of the Entomological Society of America 93:347-355.
Wahlberg N, and Zimmermann M. 2000. Pattern of phylogenetic relationships among members of the tribe Melitaeini (Lepidoptera: Nymphalidae) inferred from mitochondrial DNA sequences. Cladistics 16: 347-363.
徐堉峰. 《臺灣蝴蝶圖鑑.下【蛺蝶】》. 台灣: 晨星出版有限公司. 2013年3月10日. ISBN 9789861776682(中文(繁體)).
呂至堅、陳建仁. 《蝴蝶生活史圖鑑》. 台灣: 晨星出版有限公司. 2014年3月23日. ISBN 9789861777849(中文(繁體)).
寿建新 周尧 李宇飞. 世界蝴蝶分類名錄. 中國: 陝西科學技術出版社. 2006-04-01. ISBN 9787536936768(中文(简体)).