Brookula

Brookula
Brookula contigua
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Vetigastropoda
Superfamily:	Seguenzioidea
Family:	incertae sedis
Genus:	Brookula Iredale, 1912
Type species
Brookula stibarochila Iredale, T., 1912
Species
See text
Synonyms[1]
Benthobrookula A. H. Clarke, 1961 Brookula (Aequispirella) Finlay, 1924

Brookula is a genus of minute sea snails or micromolluscs, marine gastropod molluscs unassigned in the superfamily Seguenzioidea.^[1]

Species

Species within the genus Brookula include:^[1]

• Brookula angeli (Tenison-Woods, 1876)
• Brookula annectens Powell, 1937
• Brookula benthicola Dell, 1956
• Brookula bohni Schwabe & Engl, 2008
• Brookula brevis (d'Orbigny, 1841)
• Brookula capensis A. H. Clarke, 1961
• Brookula contigua Powell, 1940
• Brookula crebresculpta (Tate, 1899)
• Brookula decussata (Pelseneer, 1903)
• Brookula densilaminata (Verco, 1907)
• † Brookula endodonta Finlay, 1924
• Brookula finesia Laseron, 1954
• † Brookula fossilis Finlay, 1924
• † Brookula funiculata Finlay, 1924
• Brookula iki Kay, 1979
• Brookula lamonti Clarke, 1961
• Brookula megaumbilicata Absalão & Pimenta, 2005
• Brookula murawskii Fernández-Garcés, Rubio & Rolán, 2018
• Brookula prognata Finlay, 1927
• Brookula proseila Absalão & Pimenta, 2005
• † Brookula pukeuriensis Finlay, 1924
• † Brookula singletoni Chapman & Crespin, 1928
• Brookula spinulata Absalão, R.S., Miyaji & A.D. Pimenta, 2001 - Brazil
• Brookula stibarochila Iredale, 1912
• Brookula tanseimaruae Tsuchida & Hori, 1996

Species brought into synonymy

• Brookula (Aequispirella) Finlay, 1924: synonym of Aequispirella Finlay, 1924
• Brookula (Aequispirella) corula (Hutton, 1885): synonym of Brookula corula (Hutton, 1885)
• Brookula antarctica Dell, 1990: synonym of Brookula strebeli A. W. B. Powell, 1951
• Brookula argentina (Zelaya, Absalão & Pimenta, 2006): synonym of Benthobrookula argentina Zelaya, Absalão & Pimenta, 2006
• Brookula augeria Laseron, 1954: synonym of Brookula angeli (Tenison-Woods, 1877)
• Brookula bifurcata Maxwell, 1992: synonym of Aequispirella bifurcata (P. A. Maxwell, 1992)
• Brookula calypso (Melvill & Standen, 1912): synonym of Benthobrookula calypso (Melvill & Standen, 1912)
• Brookula charleenae (Schwabe & Engl, 2008): synonym of Benthobrookula charleenae (Schwabe & Engl, 2008)
• Brookula conica (R. B. Watson, 1880): synonym of Benthobrookula conica (R. B. Watson, 1880)
• Brookula consobrina May, 1923: synonym of Brookula crebresculpta (Tate, 1899)
• Brookula coronis Barnard, 1963: synonym of Imbricoscelis coronis (Barnard, 1963)
• Brookula corula (Hutton, 1885): synonym of Aequispirella corula (Hutton, 1885)
• Brookula corulum (Hutton, 1885): synonym of Aequispirella corula (Hutton, 1885)
• Brookula crassicostata (Strebel, 1908): synonym of Liotella crassicostata (Strebel, 1908)
• Brookula delli Numanami, 1996: synonym of Brookula pfefferi Powell, 1951
• Brookula enderbyensis Powell, 1931: synonym of Aequispirella enderbyensis (Powell, 1931)
• Brookula exquisita A. H. Clarke, 1961: synonym of Benthobrookula exquisita (A. H. Clarke, 1961)
• Brookula finlayi Powell, 1933: synonym of Aequispirella finlayi (Powell, 1933)
• Brookula galapagana (Dall, 1913): synonym of Vetulonia galapagana (Dall, 1913)
• Brookula gemmula (Turton, 1932): synonym of Benthobrookula gemmula (Turton, 1932)
• Brookula iredalei Finlay, 1924: synonym of Aequispirella iredalei (Finlay, 1924)
• Brookula jacksonensis Laseron, 1954: synonym of Brookula nepeanensis (Gatliff, 1906)
• Brookula kaawaensis Laws, 1940: synonym of Aequispirella kaawaensis (Laws, 1940)
• Brookula kerguelensis (Thiele, 1925): synonym of Benthobrookula kerguelensis (Thiele, 1925)
• Brookula meridionale (Melvill & Standen, 1912): synonym of Munditia meridionalis (Melvill & Standen, 1912)
• Brookula nepeanensis (Gatliff, 1906): synonym of Benthobrookula nepeanensis (Gatliff, 1906)
• Brookula obscura Laseron, 1954: synonym of Brookula angeli (Tenison-Woods, 1877)
• Brookula olearia Absalão & Pimenta, 2005: synonym of Benthobrookula olearia (Absalão & Pimenta, 2005)
• Brookula orospatia Laseron, 1954: synonym of Brookula angeli (Tenison-Woods, 1877)
• Brookula paranaensis (Zelaya, Absalão & Pimenta, 2006): synonym of Benthobrookula paranaensis Zelaya, Absalão & Pimenta, 2006
• Brookula pfefferi Powell, 1951: synonym of Benthobrookula pfefferi (A. W. B. Powell, 1951)
• Brookula polypleura (Hedley, 1904): synonym of Liotella polypleura (Hedley, 1904)
• Brookula powelli Clarke, 1961: synonym of Benthobrookula powelli (A. H. Clarke, 1961)
• Brookula rossiana Dell, 1990: synonym of Brookula pfefferi Powell, 1951
• Brookula sinusbreidensis Numanami & Okutani, 1991: synonym of Brookula pfefferi Powell, 1951
• Brookula strebeli A. W. B. Powell, 1951: synonym of Benthobrookula strebeli (A. W. B. Powell, 1951)
• Brookula tenuilirata Finlay, 1924: synonym of Aequispirella tenuilirata (Finlay, 1924)
• Brookula tumida Laseron, 1954: synonym of Brookula angeli (Tenison-Woods, 1877)
• Brookula turbinata Laseron, 1954: synonym of Brookula angeli (Tenison-Woods, 1877)

References

1. Brookula Iredale, 1912. 1 December 2025. Retrieved through: World Register of Marine Species.

• Schwabe E. & Engl W. (2008). Description of two new deep-water species of the genus Brookula Iredale, 1912 (Mollusca, Gastropoda, Trochoidea), with a revision of the genus for the Subantarctic and Arctic Sector of the Atlantic Ocean. Zootaxa, 1866: 187-204

External links

• Spencer, H.; Marshall. B. (2009). All Mollusca except Opisthobranchia. In: Gordon, D. (Ed.) (2009). New Zealand Inventory of Biodiversity. Volume One: Kingdom Animalia. 584 pp
• Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
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