List of deomyines

Eastern spiny mouse (Acomys dimidiatus)

Deomyinae is a subfamily of mammals in the rodent family Muridae, which in turn is part of the Myomorpha suborder in the order Rodentia. Members of this subfamily are called deomyines and include spiny mice and brush-furred rats. They are found in Africa, the Middle East, and Mediterranean islands, primarily in shrublands, grasslands, rocky areas, and forests, though some species can be found in deserts and savannas. They range in size from the Southern African spiny mouse, at 7 cm (3 in) plus a 6 cm (2 in) tail, to the black-clawed brush-furred rat, at 18 cm (7 in) plus a 10 cm (4 in) tail. Deomyines generally eat insects, grain, and grass, as well as other invertebrates and vegetation, frogs, and other small vertebrates. No deomyines have population estimates, but three species—the Mount Chercher brush-furred rat, Sheko Forest brush-furred rat, and Dieterlen's brush-furred mouse—are categorized as endangered, and one species—the Mount Lefo brush-furred mouse—is categorized as critically endangered.

The 55 extant species of Deomyinae are divided into 4 genera, ranging in size from 1 to 31 species. Several extinct prehistoric deomyine species have been discovered, though due to ongoing research and discoveries, the exact number and categorization are not fixed.^[1]

Conventions

IUCN Red List categories

Conservation status
EX	Extinct (0 species)
EW	Extinct in the wild (0 species)
CR	Critically endangered (1 species)
EN	Endangered (3 species)
VU	Vulnerable (2 species)
NT	Near threatened (2 species)
LC	Least concern (30 species)
Other categories
DD	Data deficient (9 species)
NE	Not evaluated (8 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the deomyine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

Deomyinae is a subfamily of the rodent family Muridae consisting of 55 extant species in 4 genera. These genera range in size from 1 to 31 species. This does not include hybrid species or extinct prehistoric species.

Subfamily Deomyinae

• Genus Acomys (spiny mice): twenty-two species
• Genus Deomys (link rat): one species
• Genus Lophuromys (brush-furred rats): thirty-one species
• Genus Uranomys (Rudd's mouse): one species

Deomyinae[2] Uranomys Acomys Deomys Lophuromys

Deomyines

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.^[3]

Genus Acomys – I. Geoffroy, 1838 – 22 species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Asia Minor spiny mouse	A. cilicicus Spitzenberger, 1978	Southern Turkey	Size: 9–13 cm (4–5 in) long, plus 9–12 cm (4–5 in) tail[4] Habitat: Shrubland and rocky areas[5] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	DD  Unknown [5]
Cairo spiny mouse	A. cahirinus (É. Geoffrey, 1803)	Northern Africa	Size: 10–12 cm (4–5 in) long, plus 9–14 cm (4–6 in) tail[4] Habitat: Shrubland and rocky areas[7] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [7]
Cape spiny mouse	A. subspinosus (Waterhouse, 1838)	Southern South Africa	Size: 7–11 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[8] Habitat: Rocky areas and shrubland[9] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [9]
Chudeau's spiny mouse	A. chudeaui Kollmann, 1911	Northwestern Africa	Size: 9–12 cm (4–5 in) long, plus 7–13 cm (3–5 in) tail[4] Habitat: Shrubland and rocky areas[7] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	NE  Unknown
Crete spiny mouse	A. minous (Bate, 1906)	Crete island in Greece	Size: 9–13 cm (4–5 in) long, plus 8–12 cm (3–5 in) tail[4] Habitat: Rocky areas and shrubland[10] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	DD  Unknown [10]
Cyprus spiny mouse	A. nesiotes (Bate, 1903)	Cyprus	Size: 9–14 cm (4–6 in) long, plus 9–12 cm (4–5 in) tail[4] Habitat: Rocky areas[11] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	DD  Unknown [11]
Eastern spiny mouse	A. dimidiatus (Cretzschmar, 1826)	Southwestern Asia and eastern Egypt	Size: 8–13 cm (3–5 in) long, plus 8–12 cm (3–5 in) tail[4] Habitat: Rocky areas, shrubland, forest, grassland, and desert[12] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [12]
Fiery spiny mouse	A. ignitus Dollman, 1910	Southern Kenya and northeastern Tanzania	Size: 9–12 cm (4–5 in) long, plus 7–9 cm (3–4 in) tail[4] Habitat: Savanna and rocky areas[13] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [13]
Golden spiny mouse	A. russatus (Wagner, 1840)	Southwestern Asia and eastern Egypt	Size: 10–12 cm (4–5 in) long, plus 6–8 cm (2–3 in) tail[8] Habitat: Rocky areas[14] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [14]
Gray spiny mouse	A. cineraceus Fitzinger & Heuglin, 1866	Eastern Africa	Size: 8–12 cm (3–5 in) long, plus 7–13 cm (3–5 in) tail[4] Habitat: Rocky areas, savanna, shrubland, and grassland[15] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [15]
Johan's spiny mouse	A. johannis Thomas, 1912	Western Africa	Size: 9–13 cm (4–5 in) long, plus 8–12 cm (3–5 in) tail[4] Habitat: Rocky areas, grassland, and shrubland[16] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [16]
Kemp's spiny mouse	A. kempi Dollman, 1911	Southwestern Asia and eastern Egypt	Size: 9–11 cm (4–4 in) long, plus 8–10 cm (3–4 in) tail[4] Habitat: Savanna and rocky areas[17] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [17]
Louise's spiny mouse	A. louisae Thomas, 1896	Eastern Africa	Size: 7–10 cm (3–4 in) long, plus 8–11 cm (3–4 in) tail[8] Habitat: Savanna, rocky areas, and grassland[18] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [18]
Mullah spiny mouse	A. mullah Thomas, 1904	Eastern Africa	Size: 10–14 cm (4–6 in) long, plus 9–12 cm (4–5 in) tail[4] Habitat: Shrubland and rocky areas[19] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [19]
Muze spiny mouse	A. muzei Verheyen, Hulselmans, Wendelen, Leirs, Corti, Backeljau, & Verheyen, 2011	Eastern Africa	Size: 7–10 cm (3–4 in) long, plus 7–11 cm (3–4 in) tail[8] Habitat: Forest[20] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [20]
Nguru spiny mouse	A. ngurui Verheyen, Hulselmans, Wendelen, Leirs, Corti, Backeljau, & Verheyen, 2011	Eastern Africa	Size: 7–11 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[8] Habitat: Forest[21] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [21]
Percival's spiny mouse	A. percivali Dollman, 1911	Eastern Africa	Size: 8–12 cm (3–5 in) long, plus 3–10 cm (1–4 in) tail[8] Habitat: Savanna, shrubland, and rocky areas[22] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [22]
Selous's spiny mouse	A. selousi de Winton, 1896	Southern Africa	Size: 7–10 cm (3–4 in) long, plus 7–10 cm (3–4 in) tail[8] Habitat: Forest, rocky areas, and shrubland[23] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [23]
Seurat's spiny mouse	A. seurati Heim de Balsac, 1936	Southern Algeria	Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail[4] Habitat: Desert and rocky areas[24] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [24]
Southern African spiny mouse	A. spinosissimus Peters, 1852	Southeastern Africa	Size: 7–10 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[8] Habitat: Rocky areas and savanna[25] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [25]
Western Saharan spiny mouse	A. airensis Thomas & Hinton, 1921	Western Africa	Size: 9–12 cm (4–5 in) long, plus 7–13 cm (3–5 in) tail[4] Habitat: Shrubland and rocky areas[26] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [26]
Wilson's spiny mouse	A. wilsoni Thomas, 1892	Eastern Africa	Size: 8–12 cm (3–5 in) long, plus 6–10 cm (2–4 in) tail[8] Habitat: Rocky areas, shrubland, and savanna[27] Diet: Omnivorous, but primarily grain, grass, and other plant material[6]	LC  Unknown [27]

Genus Deomys – Thomas, 1888 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Link rat	D. ferrugineus Thomas, 1888	Central Africa	Size: 10–15 cm (4–6 in) long, plus 15–20 cm (6–8 in) tail[28] Habitat: Inland wetlands and forest[29] Diet: Insects and other invertebrates, as well as vegetable matter[29]	LC  Unknown [29]

Genus Lophuromys – Peters, 1874 – 31 species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Angolan brush-furred rat	L. angolensis Verheyen, Dierckx, & Hulselmans, 2000	Angola and southwestern Democratic Republic of the Congo	Size: 11–16 cm (4–6 in) long, plus 5–9 cm (2–4 in) tail[8] Habitat: Grassland and forest[30] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown
Ansorge's brush-furred rat	L. ansorgei (de Winton, 1896)	Western Africa	Size: 10–17 cm (4–7 in) long, plus 4–10 cm (2–4 in) tail[8] Habitat: Grassland and forest[30] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown
Black-clawed brush-furred rat	L. melanonyx Petter, 1972	Ethiopia	Size: 12–18 cm (5–7 in) long, plus 3–10 cm (1–4 in) tail[8] Habitat: Grassland and forest[32] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	VU  Unknown [32]
Dark-colored brush-furred rat	L. cinereus Dieterlen & Gelmroth, 1974	Eastern Democratic Republic of the Congo	Size: 11–13 cm (4–5 in) long, plus 6–8 cm (2–3 in) tail[28] Habitat: Forest and grassland[33] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	DD  Unknown [33]
Dieterlen's brush-furred mouse	L. dieterleni Verheyen, Hulselmans, Colyn, & Hutterer, 1997	Western Cameroon	Size: 11–13 cm (4–5 in) long, plus 7–8 cm (3–3 in) tail[8] Habitat: Forest[34] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	EN  Unknown [34]
Dudu's brush-furred rat	L. dudui Verheyen, Hulselmans, Dierckx, & Verheyen, 2002	Northeastern Democratic Republic of the Congo	Size: 8–13 cm (3–5 in) long, plus 5–8 cm (2–3 in) tail[28] Habitat: Grassland and forest[35] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown
Ethiopian forest brush-furred rat	L. chrysopus Osgood, 1936	Ethiopia	Size: 10–13 cm (4–5 in) long, plus 7–9 cm (3–4 in) tail[28] Habitat: Forest[36] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [36]
Fire-bellied brush-furred rat	L. nudicaudus Heller, 1911	Western Africa	Size: 8–12 cm (3–5 in) long, plus 4–8 cm (2–3 in) tail[8] Habitat: Forest[37] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [37]
Gray brush-furred rat	L. aquilus (True, 1892)	Democratic Republic of the Congo	Size: 11–15 cm (4–6 in) long, plus 6–9 cm (2–4 in) tail[8] Habitat: Grassland and forest[35] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown
Hutterer's brush-furred mouse	L. huttereri Verheyen, Colyn, & Hulselmans, 1996	Democratic Republic of the Congo	Size: 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[8] Habitat: Forest[38] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [38]
Kilonzo's brush furred rat	L. kilonzoi Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti, & Verheyen, 2007	Eastern Tanzania	Size: 11–14 cm (4–6 in) long, plus 5–10 cm (2–4 in) tail[28] Habitat: Forest[39] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [39]
Machandu's brush furred rat	L. machangui Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti, & Verheyen, 2007	Southeastern Africa	Size: 10–14 cm (4–6 in) long, plus 2–9 cm (1–4 in) tail[28] Habitat: Shrubland and forest[40] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	DD  Unknown [40]
Makundi's brush-furred rat	L. makundii Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti, & Verheyen, 2007	Tanzania	Size: 11–14 cm (4–6 in) long, plus 5–9 cm (2–4 in) tail[28] Habitat: Shrubland and forest[41] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	DD  Unknown [41]
Medium-tailed brush-furred rat	L. medicaudatus Dieterlen, 1975	Central Africa	Size: 9–12 cm (4–5 in) long, plus 7–10 cm (3–4 in) tail[28] Habitat: Forest[42] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	VU  Unknown [42]
Mount Cameroon brush-furred rat	L. roseveari Verheyen, Hulselmans, Colyn, & Hutterer, 1997	Western Cameroon	Size: 10–15 cm (4–6 in) long, plus 5–8 cm (2–3 in) tail[8] Habitat: Forest, shrubland, and grassland[43] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [43]
Mount Chercher brush-furred rat	L. chercherensis Lavrenchenko, Verheyen, Verheyen, Hulselmans, & Leirs, 2007	Ethiopia	Size: 11–15 cm (4–6 in) long, plus 6–7 cm (2–3 in) tail[8] Habitat: Forest[44] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	EN  Unknown [44]
Mount Lefo brush-furred mouse	L. eisentrauti Dieterlen, 1978	Western Cameroon	Size: 8–10 cm (3–4 in) long, plus 5–6 cm (2–2 in) tail[8] Habitat: Forest[45] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	CR  Unknown [45]
North Western Rift brush-furred rat	L. menageshae Lavrenchenko, Verheyen, Verheyen, Hulselmans, & Leirs, 2007	Central Ethiopia	Size: 12–15 cm (5–6 in) long, plus 6–8 cm (2–3 in) tail[8] Habitat: Forest[46] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	DD  Unknown [46]
Rahm's brush-furred rat	L. rahmi Verheyen, 1964	Central Africa	Size: 9–12 cm (4–5 in) long, plus 4–6 cm (2–2 in) tail[8] Habitat: Forest[47] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NT  Unknown [47]
Rusty-bellied brush-furred rat	L. sikapusi (Temminck, 1853)	Western and central Africa	Size: 11–15 cm (4–6 in) long, plus 5–8 cm (2–3 in) tail[8] Habitat: Grassland and forest[30] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [30]
Sabuni's brush-furred rat	L. sabunii Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti, & Verheyen, 2007	Western Tanzania	Size: 11–15 cm (4–6 in) long, plus 3–8 cm (1–3 in) tail[28] Habitat: Shrubland and forest[48] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	DD  Unknown [48]
Sheko Forest brush-furred rat	L. pseudosikapusi Lavrenchenko, Verheyen, Verheyen, Hulselmans, & Leirs, 2007	Western Ethiopia	Size: 13–14 cm (5–6 in) long, plus 7–9 cm (3–4 in) tail[8] Habitat: Forest[49] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	EN  Unknown [49]
Short-tailed brush-furred rat	L. brevicaudus Osgood, 1936	Ethiopia	Size: 9–14 cm (4–6 in) long, plus 5–7 cm (2–3 in) tail[8] Habitat: Shrubland and grassland[50] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NT  Unknown [50]
Simien brush-furred rat	L. simensis Osgood, 1936	Northern Ethiopia	Size: 11–15 cm (4–6 in) long, plus 5–9 cm (2–4 in) tail[8] Habitat: Forest and grassland[51] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [51]
Stanley's brush-furred rat	L. stanleyi Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti, & Verheyen, 2007	Eastern Democratic Republic of the Congo and western Uganda	Size: 11–13 cm (4–5 in) long, plus 4–8 cm (2–3 in) tail[28] Habitat: Forest[52] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	DD  Unknown [52]
Thomas's Ethiopian brush-furred rat	L. brunneus Thomas, 1906	Southern Ethiopia	Size: 10–14 cm (4–6 in) long, plus 5–9 cm (2–4 in) tail[8] Habitat: Grassland and forest[35] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown
Verhagen's brush-furred rat	L. verhageni Verheyen, Hulselmans, Dierckx, & Verheyen, 2002	Northern Tanzania	Size: 11–14 cm (4–6 in) long, plus 4–9 cm (2–4 in) tail[28] Habitat: Grassland and forest[35] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown
Woosnam's brush-furred rat	L. woosnami Thomas, 1906	Central Africa	Size: 10–14 cm (4–6 in) long, plus 10–14 cm (4–6 in) tail[28] Habitat: Rocky areas, forest, and shrubland[53] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [53]
Yellow-bellied brush-furred rat	L. luteogaster Dieterlen, 1975	Eastern Democratic Republic of the Congo	Size: 9–12 cm (4–5 in) long, plus 9–12 cm (4–5 in) tail[28] Habitat: Forest[54] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [54]
Yellow-spotted brush-furred rat	L. flavopunctatus Thomas, 1888	Central and eastern Africa	Size: 12–13 cm (5–5 in) long, plus 5–7 cm (2–3 in) tail[8] Habitat: Grassland and forest[35] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	LC  Unknown [35]
Zena's brush-furred rat	L. zena Dollman, 1909	Southern Kenya	Size: 10–14 cm (4–6 in) long, plus 4–9 cm (2–4 in) tail[8] Habitat: Grassland and forest[35] Diet: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation[31]	NE  Unknown

Genus Uranomys – Dollman, 1909 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Rudd's mouse	U. ruddi Dollman, 1909	Western, central, and eastern Africa	Size: 9–12 cm (4–5 in) long, plus 6–8 cm (2–3 in) tail[28] Habitat: Savanna and shrubland[55] Diet: Insects[56]	LC  Unknown [55]

References

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23. Dando, T.; Kennerley, R. (2019). "Acomys selousi". IUCN Red List of Threatened Species. 2019 e.T47806307A47806352. doi:10.2305/IUCN.UK.2019-1.RLTS.T47806307A47806352.en.
24. Schlitter, D. (2022). "Acomys seurati". IUCN Red List of Threatened Species. 2022 e.T44939A22452894. doi:10.2305/IUCN.UK.2022-2.RLTS.T44939A22452894.en.
25. MacFadyen, D.; Medger, K. (2019). "Acomys spinosissimus". IUCN Red List of Threatened Species. 2019 e.T47805694A119392541. doi:10.2305/IUCN.UK.2019-1.RLTS.T47805694A119392541.en.
26. Schlitter, D. (2016). "Acomys airensis". IUCN Red List of Threatened Species. 2016 e.T44936A22453090. doi:10.2305/IUCN.UK.2016-2.RLTS.T44936A22453090.en.
27. Cassola, F. (2017) [amended version of 2016 assessment]. "Acomys wilsoni". IUCN Red List of Threatened Species. 2017 e.T276A113477310. doi:10.2305/IUCN.UK.2016-3.RLTS.T276A113477310.en.
28. Chernasky; Motis; Burgin, pp. 284–285
29. Schlitter, D.; Kerbis Peterhans, J. (2017) [errata version of 2016 assessment]. "Deomys ferrugineus". IUCN Red List of Threatened Species. 2016 e.T6490A115082137. doi:10.2305/IUCN.UK.2016-3.RLTS.T6490A22450212.en.
30. Cassola, F. (2017) [errata version of 2016 assessment]. "Lophuromys sikapusi". IUCN Red List of Threatened Species. 2016 e.T12356A115104961. doi:10.2305/IUCN.UK.2016-3.RLTS.T12356A22407590.en.
31. Nowak, p. 1587
32. Kennerley, R.; Lavrenchenko, L. (2016). "Lophuromys melanonyx". IUCN Red List of Threatened Species. 2016 e.T12351A22408287. doi:10.2305/IUCN.UK.2016-3.RLTS.T12351A22408287.en.
33. Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Lophuromys cinereus". IUCN Red List of Threatened Species. 2016 e.T12349A115104709. doi:10.2305/IUCN.UK.2016-3.RLTS.T12349A22408471.en.
34. Kennerley, R. (2017) [errata version of 2016 assessment]. "Lophuromys dieterleni". IUCN Red List of Threatened Species. 2016 e.T45056A115202428. doi:10.2305/IUCN.UK.2016-3.RLTS.T45056A22408180.en.
35. Cassola, F. (2016). "Lophuromys flavopunctatus". IUCN Red List of Threatened Species. 2016 e.T47990318A22407469. doi:10.2305/IUCN.UK.2016-3.RLTS.T47990318A22407469.en.
36. Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Lophuromys chrysopus". IUCN Red List of Threatened Species. 2016 e.T45057A115202576. doi:10.2305/IUCN.UK.2016-3.RLTS.T45057A22407952.en.
37. Cassola, F. (2017) [errata version of 2016 assessment]. "Lophuromys nudicaudus". IUCN Red List of Threatened Species. 2016 e.T12355A115104830. doi:10.2305/IUCN.UK.2016-3.RLTS.T12355A22407747.en.
38. Dieterlen, F. (2017). "Lophuromys huttereri". IUCN Red List of Threatened Species. 2017 e.T45059A22408026. doi:10.2305/IUCN.UK.2017-2.RLTS.T45059A22408026.en.
39. Dando, T. (2019). "Lophuromys kilonzoi". IUCN Red List of Threatened Species. 2019 e.T47998388A47998395. doi:10.2305/IUCN.UK.2019-3.RLTS.T47998388A47998395.en.
40. Verheyen, E.; Dando, T.; Kennerley, R. (2019). "Lophuromys machangui". IUCN Red List of Threatened Species. 2019 e.T47997166A47997274. doi:10.2305/IUCN.UK.2019-1.RLTS.T47997166A47997274.en.
41. Dando, T.; Verheyen, E.; Kennerley, R. (2019). "Lophuromys makundii". IUCN Red List of Threatened Species. 2019 e.T47997538A47997616. doi:10.2305/IUCN.UK.2019-1.RLTS.T47997538A47997616.en.
42. Kennerley, R. (2016). "Lophuromys medicaudatus". IUCN Red List of Threatened Species. 2016 e.T12350A22408558. doi:10.2305/IUCN.UK.2016-3.RLTS.T12350A22408558.en.
43. Dieterlen, F. (2017). "Lophuromys roseveari". IUCN Red List of Threatened Species. 2017 e.T45055A22408108. doi:10.2305/IUCN.UK.2017-2.RLTS.T45055A22408108.en.
44. Dando, T.; Kennerley, R.; Lavrenchenko, L.; Verheyen, E. (2020) [amended version of 2019 assessment]. "Lophuromys chercherensis". IUCN Red List of Threatened Species. 2020 e.T47992974A166612346. doi:10.2305/IUCN.UK.2020-1.RLTS.T47992974A166612346.en.
45. Hoffmann, M.; Kennerley, R. (2016). "Lophuromys eisentrauti". IUCN Red List of Threatened Species. 2016 e.T136619A22408665. doi:10.2305/IUCN.UK.2016-3.RLTS.T136619A22408665.en.
46. Dando, T.; Kennerley, R.; Verheyen, E.; Lavrenchenko, L. (2019). "Lophuromys menageshae". IUCN Red List of Threatened Species. 2019 e.T47992929A47992934. doi:10.2305/IUCN.UK.2019-1.RLTS.T47992929A47992934.en.
47. Kennerley, R. (2016). "Lophuromys rahmi". IUCN Red List of Threatened Species. 2016 e.T12352A22408380. doi:10.2305/IUCN.UK.2016-3.RLTS.T12352A22408380.en.
48. Dando, T.; Verheyen, E.; Kennerley, R. (2019). "Lophuromys sabunii". IUCN Red List of Threatened Species. 2019 e.T47997414A47997488. doi:10.2305/IUCN.UK.2019-1.RLTS.T47997414A47997488.en.
49. Dando, T.; Kennerley, R.; Lavrenchenko, L.; Verheyen, E. (2018). "Lophuromys pseudosikapusi". IUCN Red List of Threatened Species. 2018 e.T47992992A47993010. doi:10.2305/IUCN.UK.2018-1.RLTS.T47992992A47993010.en.
50. Verheyen, E.; Lavrenchenko, L.; Dando, T. (2020). "Lophuromys brevicaudus". IUCN Red List of Threatened Species. 2020 e.T45058A22407828. doi:10.2305/IUCN.UK.2020-2.RLTS.T45058A22407828.en.
51. Lavrenchenko, L.; Verheyen, E.; Kennerley, R.; Dando, T. (2019). "Lophuromys simensis". IUCN Red List of Threatened Species. 2019 e.T47992103A47992768. doi:10.2305/IUCN.UK.2019-1.RLTS.T47992103A47992768.en.
52. Dando, T.; Verheyen, E.; Kennerley, R. (2019). "Lophuromys stanleyi". IUCN Red List of Threatened Species. 2019 e.T47996434A47997111. doi:10.2305/IUCN.UK.2019-1.RLTS.T47996434A47997111.en.
53. Cassola, F. (2016). "Lophuromys woosnami". IUCN Red List of Threatened Species. 2016 e.T12357A22407309. doi:10.2305/IUCN.UK.2016-2.RLTS.T12357A22407309.en.
54. Cassola, F. (2016). "Lophuromys luteogaster". IUCN Red List of Threatened Species. 2016 e.T12354A22407410. doi:10.2305/IUCN.UK.2016-2.RLTS.T12354A22407410.en.
55. Granjon, L. (2017). "Uranomys ruddi". IUCN Red List of Threatened Species. 2017 e.T22771A22400326. doi:10.2305/IUCN.UK.2017-2.RLTS.T22771A22400326.en.
56. Nowak, p. 1593

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