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↑Pearson, N. E. 1924. The fishes of the eastern slope of the Andes. I. The fishes of the Rio Beni basin, Bolivia, collected by the Mulford expedition. Ind. Univ. Studies v. 11 (núm. 64): 1-83, Pls. 1-12.
↑Messner, E. 1962. Lista de los peces Tetragonopterinae (fam. Characidae) del Uruguay. Bol. Asoc. Lat. Amer. Ictiol. Herpet. v. 2 (núm. 1): 4-5.
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