↑Breder, C. M.; Rosen, D. E. Modes of Reproduction in Fishes (en anglès). Museu Americà d'Història Natural, 1966.
↑Crivelli, A.J. 1996. The freshwater fish endemic to the Mediterranean region. An action plan for their conservation. Tour du Valat Publication, 171 p.
↑Kottelat, M. i J. Freyhof 2007. Handbook of European freshwater fishes. Publications Kottelat, Cornol, Suïssa. 646 p.
Borbori, D.C., P.S. Economidis i E.G. Maurakis 2001. Freshwater fish habitat science and management in Greece. Aquatic Ecosystem Health and Management 4: 381-391.
Economidis, P.S. 1991. Check list of freshwater fishes of Greece (recent status of threats and protection). Hellenic Society for the Protection of Nature, Special publication, 48 p.
Economidis, P.S., E. Dimitriou, R. Pagoni, E. Michaloudi i L. Natsis 2000. Introduced and translocated fish species in the inland waters of Greece. Fish. Manage. Ecol. 7:239-250.
Eschmeyer, W. N. Genera of Recent Fishes (en anglès). iii + 697. Acadèmia de les Ciències de Califòrnia, 1990. ISBN 0-940228-23-8..