Coloma, Duellman, Almendáriz, Ron, Terán-Valdez, Guaysamin, 2010 : Five new (extinct?) species of Atelopus (Anura: Bufonidae) from Andean Colombia, Ecuador, and Peru. Zootaxa, no 2574, p. 1–54.
↑Coloma, Duellman, Almendáriz, Ron, Terán-Valdez, Guaysamin, 2010 : Five new (extinct?) species of Atelopus (Anura: Bufonidae) from Andean Colombia, Ecuador, and Peru. Zootaxa, no 2574, p. 1–54.