Dyschirius

Dyschirius
Dyschirius intermedius
Nomenklatur
Dyschirius Bonelli, 1810
Klassifikasjon
Rike		Dyreriket
Rekke		Leddyr
Klasse		Insekter
Orden		Biller
Familie		Løpebiller
Underfamilie		Graveløpere
Økologi
Antall arter:		ca. 290  ? 11 i Norge
Habitat:		på land
Utbredelse:		kosmopolittisk
Inndelt i

Dyschirius er en slekt av billene i delgruppen (underfamilien) graveløpere (Scaritinae) i familiegruppen løpebiller.

Utseende

Små (2,5-7,5 millimeter), avlange, blanke, mørke løpebiller med kraftige bein. Pronotum og dekkvingene er rundet hver for seg, overflatisk kan de minne vagt om maur. Antennene er perlekjede-formede og forholdsvis korte. Hodet er forholdsvis lite med middelsstore fasettøyne og kjever. Pronotum er rundet og hvelvet, dekkvingene rundede. Beina er utviklet til gravebein.

Levevis

Disse billene graver i bakken og lever for det meste underjordisk. Flere av artene lever på havstrender, der de gjerne graver seg ned i sand eller mudder ved flo sjø. De er rovdyr som jakter på små leddyr.

Utbredelse

Slekten forekommer i alle verdensdeler bortsett fra Australia (og Antarktis). De fleste av artene lever Palearktis.

Systematisk inndeling

Oversikten har med de europeiske artene.^[1] Systematikken følger Carabidae of the World.^[2]

Treliste

• ordenen biller, Coleoptera Linnaeus, 1758
  • underordenen Adephaga Schellenberg, 1806
    • overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
      • familien løpebiller, Carabidae Latreille, 1802
        • underfamilie graveløpere Scaritinae Bonelli, 1810
          • stammen Dyschiriini W. Kolbe, 1880
            • slekten Dyschirius Bonelli, 1810 - arter som forekommer i Norge:
              • Dyschirius angustatus (Ahrens, 1830)
              • Dyschirius armatus Wollaston 1864
              • Dyschirius caspius Putzeys 1866
              • Dyschirius digitatus (Dejean 1825)
              • Dyschirius humeratus Chaudoir 1850
              • Dyschirius latipennis Seidlitz 1867
              • Dyschirius numidicus Putzeys 1846
              • Dyschirius obscurus (Gyllenhal, 1827)
              • Dyschirius thoracicus (P. Rossi, 1790)
              • Dyschirius zimini Znojko 1928

Liste over arter som ikke forekommer i Norge

• Dyschirius abbreviatus Putzeys, 1846 ^{i c g b}
• Dyschirius abditus Fedorenko, 1993 ^{c g}
• Dyschirius addisabeba (Bulirsch, 2006) ^{c g}
• Dyschirius aenea Boheman, 1849
• Dyschirius aeneo Boheman, 1849 ^c
• Dyschirius aeneolus Leconte, 1850
• Dyschirius aeneus (Dejean, 1825) ^{c g}
• Dyschirius affinis Fall, 1901 ^{i c g}
• Dyschirius afghanus Jedlicka, 1967 ^{c g}
• Dyschirius agnatus Motschulsky, 1844 ^{c g}
• Dyschirius aida Schatzmayr, 1936 ^{c g}
• Dyschirius alajensis Znojko, 1930 ^{c g}
• Dyschirius alticola Lindroth, 1961 ^{i c g}
• Dyschirius amazonicus Fedorenko, 1991 ^{c g}
• Dyschirius ambiguus Fedorenko, 1994 ^{c g}
• Dyschirius amphibolus J. Muller, 1922
• Dyschirius amurensis Fedorenko, 1991 ^{c g}
• Dyschirius analis Leconte, 1852
• Dyschirius angolensis (Fedorenko, 2000) ^{c g}
• Dyschirius angusticollis Putzeys, 1867 ^{c g}
• Dyschirius anichtchenkoi (Bulirsch, 2015) ^{c g}
• Dyschirius apicalis Putzeys, 1846 ^{c g}
• Dyschirius apicius Gistel, 1857 ^c
• Dyschirius aratus Leconte, 1852
• Dyschirius arcanus Bruneau de Miré, 1952 ^{c g}
• Dyschirius arcifer Znojko, 1928 ^{c g}
• Dyschirius arnoldii Gryuntal, 1984 ^{c g}
• Dyschirius asper Andrewes, 1929 ^{c g}
• Dyschirius assegaaicus (Fedorenko, 2000) ^{c g}
• Dyschirius auriculatus Wollaston, 1867 ^{c g}
• Dyschirius bacillus Schaum, 1857 ^{c g}
• Dyschirius baehri (Bulirsch, 2006) ^{c g}
• Dyschirius baenningeri (Fedorenko, 2004) ^{c g}
• Dyschirius baicalensis Motschulsky, 1844 ^{c g}
• Dyschirius bechynei Kult, 1949 ^{c g}
• Dyschirius becvari (Bulirsch, 2006) ^{c g}
• Dyschirius beludscha Tschitscherine, 1904
• Dyschirius beludsha Tschitscherine, 1904 ^{c g}
• Dyschirius benedikti Bulirsch, 1995 ^{c g}
• Dyschirius bengalensis Andrewes, 1929 ^{c g}
• Dyschirius beydagensis Jeanne, 1996 ^{c g}
• Dyschirius bifrons Andrewes, 1929 ^{c g}
• Dyschirius binodosus Putzeys, 1878 ^{c g}
• Dyschirius boliviensis (Bulirsch, 2015) ^{c g}
• Dyschirius bonellii Putzeys, 1846 ^{c g}
• Dyschirius bousqueti (Bulirsch, 2006) ^{c g}
• Dyschirius braziliensis Fedorenko, 1999
• Dyschirius breviphthalmus Balkenohl & Lompe, 2003 ^{c g}
• Dyschirius brevispinus Leconte, 1878
• Dyschirius bruchi (Kult, 1950) ^{c g}
• Dyschirius bryanti (Kult, 1950) ^{c g}
• Dyschirius buglanensis Bulirsch, 1996 ^{c g}
• Dyschirius campicola Lindroth, 1961 ^{i c g b}
• Dyschirius cariniceps Baudi di Selve, 1864 ^{c g}
• Dyschirius carri Bousquet, 1996 ^{c g}
• Dyschirius carrorum Bousquet, 1997 ⁱ
• Dyschirius cerberus Larson, 1968 ^{i c g}
• Dyschirius chalceus Erichson, 1837 ^{c g}
• Dyschirius chalybeus Putzeys, 1846 ^{c g}
• Dyschirius championi Andrewes, 1929 ^{c g}
• Dyschirius changlingensis Li, 1992 ^{c g}
• Dyschirius cheloscelis Bates, 1873 ^{c g}
• Dyschirius chiricahuae Dajoz, 2004
• Dyschirius clorinda Bulirsch, 2009
• Dyschirius clypeatus Putzeys, 1867 ^{c g}
• Dyschirius comatus Bousquet, 1988 ^{i c g}
• Dyschirius compactus Lindroth, 1961 ^{i c g}
• Dyschirius consobrinus Leconte, 1852
• Dyschirius constrictus Andrewes, 1929 ^{c g}
• Dyschirius contortus (Fedorenko, 1997) ^{c g}
• Dyschirius crenulatus Putzeys, 1867 ^{c g}
• Dyschirius criddlei Fall, 1925 ^{i c g}
• Dyschirius crinifer Balkenohl, 1993 ^{c g}
• Dyschirius curvispinus Putzeys, 1846 ^{i c g}
• Dyschirius cylindricus (Dejean, 1825) ^{c g}
• Dyschirius darlingtoni Kult, 1950 ^{c g}
• Dyschirius darwini (Kult, 1950) ^{c g}
• Dyschirius dejeanii Putzeys, 1846 ^{i c g b}
• Dyschirius devroeyianus Burgeon, 1935 ^{c g}
• Dyschirius dimidiatus Chaudoir, 1846 ^{c g}
• Dyschirius disjunctus Andrewes, 1929 ^{c g}
• Dyschirius dispar Péringuey, 1896 ^{c g}
• Dyschirius distinctus Motschulsky, 1850 ^c
• Dyschirius dostali (Bulirsch & Fedorenko, 2007) ^{c g}
• Dyschirius ecuadorensis (Bulirsch, 2006) ^{c g}
• Dyschirius edentulus Putzeys, 1846 ^{i c g}
• Dyschirius eduardinus Burgeon, 1935 ^{c g}
• Dyschirius erwini Bulirsch, 2009
• Dyschirius erythrocerus Leconte, 1857 ^{i c g b}
• Dyschirius euphraticus Putzeys, 1846 ^{c g}
• Dyschirius euxinus Znojko, 1927 ^{c g}
• Dyschirius exaratus Putzeys, 1867 ^{c g}
• Dyschirius exochus Whitehead, 1970 ^{i c g}
• Dyschirius extensus Putzeys, 1846 ^{c g}
• Dyschirius fabbrii (Bulirsch & Magrini, 2006) ^{c g}
• Dyschirius facchinii Bulirsch, 2009
• Dyschirius fassatii Kult, 1949 ^{c g}
• Dyschirius fedorenkoi (Bulirsch, 2006) ^{c g}
• Dyschirius ferganensis Znojko, 1930 ^{c g}
• Dyschirius ferrugineus Bousquet, 1988 ^{i c g}
• Dyschirius fianarensis Bulirsch, 2006 ^{c g}
• Dyschirius filiformis Leconte, 1857
• Dyschirius flavicornis Péringuey, 1896 ^{c g}
• Dyschirius fleischeri Sainte-Claire Deville, 1904 ^{c g}
• Dyschirius fossifrons Putzeys, 1867 ^{c g}
• Dyschirius franzi Kult, 1954 ^{c g}
• Dyschirius freyi Jedlicka, 1958 ^{c g}
• Dyschirius fulgidus Motschulsky, 1850 ^{c g}
• Dyschirius fulvipes (Dejean, 1825) ^{c g}
• Dyschirius fulvus (Fedorenko, 2000) ^{c g}
• Dyschirius fusus Putzeys, 1878 ^{c g}
• Dyschirius genieri Bulirsch, 2009
• Dyschirius gerardi Burgeon, 1935 ^{c g}
• Dyschirius gibbipennis Leconte, 1857
• Dyschirius girardi Kult, 1949 ^{c g}
• Dyschirius globosus Herbst, 1784
• Dyschirius globulosus (Say, 1823) ^{i c g b}
• Dyschirius gracilis (Heer, 1837) ^{c g}
• Dyschirius guatemalenus Bates, 1881 ^{c g}
• Dyschirius haemorrhoidalis (Dejean, 1831) ^{i c g b}
• Dyschirius hajeki Bulirsch, 2009
• Dyschirius hassei Kult, 1954
• Dyschirius hessei Kult, 1954 ^{c g}
• Dyschirius heydeni A.Fleischer, 1899 ^{c g}
• Dyschirius hiemalis Bousquet, 1987 ^{i g b}
• Dyschirius himalaicus (Fedorenko, 1997) ^{c g}
• Dyschirius hingstoni Andrewes, 1929 ^{c g}
• Dyschirius hiogoensis Bates, 1873 ^{c g}
• Dyschirius hipponensis Pic, 1894 ^{c g}
• Dyschirius hoberlandti Kult, 1954 ^{c g}
• Dyschirius humiolcus Chaudoir, 1850 ^{c g}
• Dyschirius importunoides Jeanne, 1996 ^{c g}
• Dyschirius importunus Schaum, 1857 ^{c g}
• Dyschirius impressifrons Fedorenko, 1993 ^{c g}
• Dyschirius impressus Putzeys, 1846 ^{c g}
• Dyschirius impunctipennis Dawson, 1854 ^{c g}
• Dyschirius insularis (Bulirsch & Fedorenko, 2013) ^{c g}
• Dyschirius integer Leconte, 1852
• Dyschirius interior Fall, 1922 ^{i c g b}
• Dyschirius intermedius Putzeys, 1846 ^{c g}
• Dyschirius intricatus (Fedorenko, 2000) ^{c g}
• Dyschirius jedlickai Kult, 1940 ^{c g}
• Dyschirius jelineki Bulirsch, 2009
• Dyschirius jindrai Fedorenko, 2004
• Dyschirius jordanicus Fedorenko, 1996
• Dyschirius kabakovi (Fedorenko, 1996) ^{c g}
• Dyschirius kadleci Bulirsch, 2009
• Dyschirius kaliki Kult, 1949 ^{c g}
• Dyschirius katanganus Burgeon, 1935 ^{c g}
• Dyschirius kirghizicus Fedorenko, 1994 ^{c g}
• Dyschirius kirschenhoferi (Bulirsch, 2012) ^{c g}
• Dyschirius kryzhanovskii Gryuntal, 1984 ^{c g}
• Dyschirius lacustris Andrewes, 1929 ^{c g}
• Dyschirius laevifasciatus G.Horn, 1878
• Dyschirius laeviusculus Putzeys, 1846 ^{c g}
• Dyschirius lambertoni Vuillet, 1910 ^{c g}
• Dyschirius larochellei Bousquet, 1988 ^{i c g b}
• Dyschirius lgockii A.Fleischer, 1912 ^{c g}
• Dyschirius limpopo (Fedorenko, 2000) ^{c g}
• Dyschirius longicollis Motschulsky, 1844 ^{c g}
• Dyschirius longipennis Putzeys, 1867 ^{c g}
• Dyschirius longulus Leconte, 1850 ^{i c g b}
• Dyschirius luticola Chaudoir, 1850 ^{c g}
• Dyschirius luzonicus Kult, 1949 ^{c g}
• Dyschirius macroderus Chaudoir, 1850 ^{c g}
• Dyschirius macrophthalmus (Fedorenko, 1999) ^{c g}
• Dyschirius mafuga (Bulirsch, 2006) ^{c g}
• Dyschirius magrinii (Bulirsch, 2012) ^{c g}
• Dyschirius mahratta Andrewes, 1929 ^{c g}
• Dyschirius malawicus (Bulirsch, 2006) ^{c g}
• Dyschirius marani Kult, 1949 ^{c g}
• Dyschirius matisi Lafer, 1989 ^{c g}
• Dyschirius melancholicus Putzeys, 1867 ^{i c g}
• Dyschirius mesopotamicus J. Muller, 1922
• Dyschirius microthorax Motschulsky, 1844 ^{c g}
• Dyschirius milloti Jeannel, 1949 ^{c g}
• Dyschirius minarum Putzeys, 1867 ^{c g}
• Dyschirius minutus (Dejean, 1825) ^{c g}
• Dyschirius montanus Leconte, 1879
• Dyschirius moraveci (Bulirsch, 2006) ^{c g}
• Dyschirius morio Putzeys, 1867 ^{c g}
• Dyschirius mortchaensis Bruneau de Miré, 1952 ^{c g}
• Dyschirius natalensis (Fedorenko, 1997) ^{c g}
• Dyschirius neoteutonus Fedorenko, 1991 ^{c g}
• Dyschirius neresheimeri Wagner, 1915
• Dyschirius nianus Fedorenko, 1993 ^{c g}
• Dyschirius nigricornis Motschulsky, 1844 ^{i c g}
• Dyschirius nitens Putzeys, 1878 ^{c g}
• Dyschirius nitidus (Dejean, 1825) ^{c g}
• Dyschirius ogloblini (Kult, 1950) ^{c g}
• Dyschirius opistholius Alluaud, 1936 ^{c g}
• Dyschirius ordinatus Bates, 1873 ^{c g}
• Dyschirius orientalis Putzeys, 1867 ^{c g}
• Dyschirius owen Dajoz, 2004
• Dyschirius pacificus Lindroth, 1961 ^{i c g}
• Dyschirius paleki Kult, 1949 ^{c g}
• Dyschirius pallipennis (Say, 1823) ^{i c g b}
• Dyschirius pampicola Putzeys, 1867 ^{c g}
• Dyschirius parallelus Motschulsky, 1844 ^{c g}
• Dyschirius parvulus Péringuey, 1896 ^{c g}
• Dyschirius patruelis Leconte, 1852
• Dyschirius paucipunctus Andrewes, 1929 ^{c g}
• Dyschirius pauxillus Wollaston, 1864 ^{c g}
• Dyschirius peringueyi Kult, 1954 ^{c g}
• Dyschirius persicus Fedorenko, 1994 ^{c g}
• Dyschirius peruanus Fedorenko, 1991 ^{c g}
• Dyschirius perversus Fall, 1922 ^{i c g}
• Dyschirius peyrierasi Basilewsky, 1976 ^{c g}
• Dyschirius pfefferi Kult, 1949 ^{c g}
• Dyschirius pilosus Leconte, 1857 ^{i c g b}
• Dyschirius planatus Lindroth, 1961 ^{i c g b}
• Dyschirius planiusculus Putzeys, 1867 ^{c g}
• Dyschirius politus (Dejean, 1825) ^{i c g b}
• Dyschirius pumilus (Dejean, 1825) ^{i c g b}
• Dyschirius punctatus (Dejean, 1825) ^{c g}
• Dyschirius pusillus (Dejean, 1825) ^{c g}
• Dyschirius quadrimaculatus Lindroth, 1961 ^{i c g b}
• Dyschirius recurvus Putzeys, 1867 ^{c g}
• Dyschirius reitteri Kult, 1949 ^{c g}
• Dyschirius remotepunctatus Putzeys, 1867 ^c
• Dyschirius roubali Maran, 1938 ^{c g}
• Dyschirius rufimanus A.Fleischer, 1898 ^{c g}
• Dyschirius rufipes (Dejean, 1825) ^{c g}
• Dyschirius rugifer Putzeys, 1878 ^{c g}
• Dyschirius ruthmuellerae Bulirsch, 2009
• Dyschirius sabahensis Bulirsch, 2009
• Dyschirius safraneki (Bulirsch & Fedorenko, 2013) ^{c g}
• Dyschirius salinus Schaum, 1843 ^{c g}
• Dyschirius salivagans Leconte, 1875
• Dyschirius saudiarabicus Balkenohl, 1994 ^{c g}
• Dyschirius schaumii Putzeys, 1867 ^{c g}
• Dyschirius scriptifrons A.Fleischer, 1898 ^{c g}
• Dyschirius sculptus Bousquet, 1988 ^{i c g}
• Dyschirius sellatus Leconte, 1857
• Dyschirius selvas (Fedorenko, 1999) ^{c g}
• Dyschirius semistriatus (Dejean, 1825) ^{c g}
• Dyschirius senegalensis Bruneau de Miré, 1952 ^{c g}
• Dyschirius setosus Leconte, 1857
• Dyschirius sevanensis Iablokoff-Khnzorian, 1962 ^{c g}
• Dyschirius sextoni Bousquet, 1987 ^{i c g}
• Dyschirius shaumii Putzeys, 1866
• Dyschirius similatus Betta, 1847 ^c
• Dyschirius sjoestedti G.Muller, 1935
• Dyschirius sjostedti G.Müller, 1935 ^{c g}
• Dyschirius smetanai (Bulirsch, 2011) ^{c g}
• Dyschirius smyrnensis (Fedorenko, 1996) ^{c g}
• Dyschirius snizeki (Bulirsch, 2011) ^{c g}
• Dyschirius soda Dajoz, 2004
• Dyschirius sonamargensis Balkenohl, 1994 ^{c g}
• Dyschirius speculifer Andrewes, 1929 ^{c g}
• Dyschirius sphaericollis (Say, 1823) ^{i c g b}
• Dyschirius sphaerulifer Bates, 1873 ^{c g}
• Dyschirius stellula Andrewes, 1936 ^{c g}
• Dyschirius steno Bates, 1873 ^{c g}
• Dyschirius stenoderus Putzeys, 1873 ^{c g}
• Dyschirius strumosus Erichson, 1837 ^{c g}
• Dyschirius subarcticus Lindroth, 1961 ^{i c g}
• Dyschirius subcylindricus Motschulsky, 1849
• Dyschirius sublaevis Putzeys, 1846 ^{i c g b}
• Dyschirius substriatus (Duftschmid, 1812) ^{c g}
• Dyschirius syriacus Putzeys, 1867 ^{c g}
• Dyschirius szeli Bulirsch, 2006 ^{c g}
• Dyschirius tamil Andrewes, 1929 ^{c g}
• Dyschirius tardipes Gistel, 1857 ^c
• Dyschirius tenuescens Andrewes, 1929 ^{c g}
• Dyschirius tenuispinus Lindroth, 1961 ^{i c g b}
• Dyschirius terminatus Leconte, 1848
• Dyschirius thailandicus (Fedorenko, 1996) ^{c g}
• Dyschirius timidus Lindroth, 1961 ^{i c g}
• Dyschirius tonkinensis (Bulirsch & Fedorenko, 2013) ^{c g}
• Dyschirius tricuspis Andrewes, 1929 ^{c g}
• Dyschirius tridentatus Leconte, 1852
• Dyschirius tristis Stephens, 1827 ^{c g}
• Dyschirius truncatus Leconte, 1857
• Dyschirius unipunctatus Fall, 1901 ^{i c g}
• Dyschirius ussuriensis Fedorenko, 1991 ^{c g}
• Dyschirius vadoni Jeannel, 1946 ^{c g}
• Dyschirius vanhillei Basilewsky, 1962
• Dyschirius varidens Fall, 1910 ^{i c g b}
• Dyschirius verticalis Putzeys, 1878 ^{c g}
• Dyschirius vietnamicus (Fedorenko, 2000) ^{c g}
• Dyschirius wayah Dajoz, 2005
• Dyschirius weyrauchi Kult, 1950 ^{c g}
• Dyschirius yezoensis Bates, 1883 ^{c g}
• Dyschirius zambesiensis (Fedorenko, 2000) ^{c g}
• Dyschirius zanzibaricus Chaudoir, 1878 ^{c g}

Referanser

1. Dyschirius - Fauna Europaea søkedato: januar 2009
2. «Carabidae of the World». Arkivert fra originalen 16. desember 2008. Besøkt 2. januar 2009. 

Litteratur

• Bertram G. Rye. 1908. Biller I. Løpebiller. Danmarks Fauna Bind 3. G.E.C. GAD, København.

Eksterne lenker

• (en) Dyschirius – oversikt og omtale av artene i WORMS-databasen
• (en) Dyschirius i Encyclopedia of Life
• (en) Dyschirius i Global Biodiversity Information Facility
• (no) Dyschirius hos Artsdatabanken
• (sv) Dyschirius hos Dyntaxa
• (en) Dyschirius hos Fauna Europaea
• (en) Dyschirius hos Fossilworks
• (en) Dyschirius hos ITIS
• (en) Dyschirius hos NCBI
• (en) Kategori:Dyschirius – bilder, video eller lyd på Wikimedia Commons
• Dyschirius – detaljert informasjon på Wikispecies
• Dyschirius – Database Carabidae of the World – engelskspråklig
• Dyschirius - Fauna Europaea - engelskspråklig
• Norsk Entomologisk forening

Autoritetsdata