Portal:Devonian

The Devonian Portal

A map of Earth as it appeared 390 million years ago during the Middle Devonian Epoch

The Devonian (/dəˈvoʊni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.62 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.86 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician Period.

Fishes, especially jawed fish, reached substantial diversity during this time, leading the Devonian to be called the Age of Fishes. The armored placoderms began dominating almost every known aquatic environment. In the oceans, cartilaginous fishes such as primitive sharks became more numerous than in the Silurian and Late Ordovician. Tetrapodomorphs, which include the ancestors of all four-limbed vertebrates (i.e. tetrapods), began diverging from freshwater lobe-finned fish as their more robust and muscled pectoral and pelvic fins gradually evolved into forelimbs and hindlimbs, though they were not fully established for life on land until the Late Carboniferous. (Full article...)

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Selected Devonian Article

The Acadian orogeny is a long-lasting mountain building event which began in the Middle Devonian, reaching a climax in the Late Devonian. It was active for approximately 50 million years, beginning roughly around 375 million years ago (Ma), with deformational, plutonic, and metamorphic events extending into the early Mississippian. The Acadian orogeny is the third of the four orogenies that formed the Appalachian Mountains and subsequent basin. The preceding orogenies consisted of the Grenville and Taconic orogenies, which followed a rift/drift stage in the Neoproterozoic. The Acadian orogeny involved the collision of a series of Avalonian continental fragments with the Laurasian continent. Geographically, the Acadian orogeny extended from the Canadian Maritime provinces migrating in a southwesterly direction toward Alabama. However, the northern Appalachian region, from New England northeastward into Gaspé region of Canada, was the most greatly affected region by the collision.

It was roughly contemporaneous with the Bretonic phase of the Variscan orogeny of Laurussia, with metamorphic events in southwestern Texas and northern Mexico, and with the Antler orogeny of the Great Basin. (Full article...)

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Selected Devonian land plant article

Archaeopteris fossil leaves

The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the spermatophytes, ancestral to both gymnosperms and angiosperms (flowering plants). They have been treated formally at the rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as Protopteridium, in which the vegetative organs consisted of relatively loose clusters of axes. Tetraxylopteris is another example of a genus lacking leaves. In more advanced aneurophytaleans such as Aneurophyton these vegetative organs started to look rather more like fronds, and eventually during Late Devonian times the aneurophytaleans are presumed to have given rise to the pteridosperm order, the Lyginopteridales. In Late Devonian times, another group of progymnosperms gave rise to the first really large trees known as Archaeopteris. The latest surviving group of progymnosperms is the Noeggerathiales, which persisted until the end of the Permian.

Other characteristics:

Vascular cambium with unlimited growth potential is present as well as xylem and phloem.
Ancestors of the earliest seed plants as well as the first true trees.

Strong monopodial growth is exhibited. (Full article...)

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The Traverse Group is a geologic group in Michigan, Indiana and Ohio comprising middle Devonian limestones with calcareous shale components. Its marine fossils notably include Michigan's state stone, the Petoskey stone (the extinct coral Hexagonaria percarinata), among other corals and records of ancient marine life. A range of trilobites has also been found in the Traverse Group.

The Traverse Group outcrops in Emmet and Charlevoix counties along the northwestern shore of Michigan's lower peninsula. Its formations are Gravel Point, Charlevoix Limestone, Petoskey, and Whiskey Creek. The Gravel Point Formation consists of a lithographic gray to brown limestone with shale beds up to 0.5 meters thick; it also includes chert nodules and bioherms (fossilized reef mounds). The Charlevoix Limestone is a mildly argillaceous limestone with interbedded coquina. The Petoskey Formation is an arenaceous limestone named for its locale (Petoskey, Michigan), and contains the eponymous Petoskey stones. The Whiskey Creek Formation is a limestone.

The Traverse Group formed as a shallow carbonate shelf during the Devonian period (~419 to 359 Ma), when the most recent supercontinent, Pangea, was just beginning to take shape. (Full article...)

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Selected Devonian fish article

Fossil of Bothriolepis panderi, an antiarch placoderm showing its caliper-like pectoral fins

Placoderms (from Ancient Greek πλάξ [plax, plakos] 'plate' and δέρμα [derma] 'skin') are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates (hence the name), and the rest of the body was scaled or naked depending on the species.

Placoderms were among the first jawed fish (their jaws likely evolved from the first pair of gill arches), as well as the first vertebrates to have true teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, Incisoscutum, Materpiscis and Austroptyctodus, represent the oldest known examples of live birth.

Placoderms are thought to be paraphyletic, consisting of several distinct outgroups or sister taxa to all living jawed vertebrates, which originated among their ranks. In contrast, one 2016 analysis concluded that Placodermi is likely monophyletic. (Full article...)

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Selected Devonian invertebrate

Palaeocharinus rhyniensis life restoration at MUSE - Science Museum in Trento

Palaeocharinus is a genus of extinct trigonotarbid arachnids known from the Devonian of western Europe. The genus was first found and described in the Rhynie chert in the 1920s by Arthur Stanley Hirst and S. Maulik. The family to which the genus belongs may be paraphyletic. (Full article...)