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↑Boulenger, G. A. 1911. Descriptions of three new characinid fishes from south-western Colombia. Annals and Magazine of Natural History (Series 8) v. 7 (núm. 38): 212-213.
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↑Eigenmann, C. H. 1922. The fishes of western South America, Part I. The fresh-water fishes of northwestern South America, including Colombia, Panama, and the Pacific slopes of Ecuador and Peru, together with an appendix upon the fishes of the Rio Meta in Colombia. Memoirs of the Carnegie Museum v. 9 (núm. 1): 1-346, Pls. 1-38.
↑Günther, A. 1860. Second list of cold-blooded vertebrata collected by Mr. Fraser in the Andes of western Ecuador. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London 1859 (pt 3): 402-420.
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