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↑Duff, R. Joel; Villarreal, Juan Carlos; Cargill, D. Christine; Renzaglia, Karen S. (2007). «Progress and challenges toward a phylogeny and classification of the hornworts». The Bryologist. 110 (2): 214–243. doi:10.1639/0007-2745(2007)110[214:PACTDA]2.0.CO;2