↑ 1,01,1Cuvier, G. & A. Valenciennes, 1850. Histoire naturelle des poissons. Tome vingt-deuxième. Suite du livre vingt-deuxième. Suite de la famille des Salmonoïdes. Table générale de l'Histoire Naturelle des Poissons (pp. 1-91). Hist. Nat. Poiss. v. 22: i-xx + 1 p. + 1-532 + 1-91, Pls. 634-650.
↑Regan, C. T., 1903. Descriptions of new South-American fishes in the collection of the British Museum. Annals and Magazine of Natural History (Series 7) v. 12 (núm. 72): 621-630.
↑Fowler, H. W., 1911. New fresh-water fishes from western Ecuador. Proc. Acad. Nat. Sci. Phila. v. 63: 493-520.
↑Bussing, W. A., 1967. New species and new records of Costa Rican freshwater fishes with a tentative list of species. Revista de Biología Tropical v. 14 (núm. 2): 205-249.
↑Boulenger, G. A., 1887. An account of the fishes collected by Mr. C. Buckley in eastern Ecuador. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London 1887 (pt 2): 274-283, Pls. 20-24.
↑Boulenger, G. A., 1899. Viaggio del Dott. Enrico Festa nel Darien e regioni vicine. Poissons de l'Amérique Centrale. Teleostei. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino v. 14 (no. 346): 1-4.
↑Gill, T. N., 1877. Notes on fishes from the Isthmus of Panama, collected by Dr. J. F. Bransford, U. S. N. Proceedings of the Academy of Natural Sciences of Philadelphia v. 28 (for 1876): 335-339.
↑Regan, C. T., 1903. Descriptions of new South-American fishes in the collection of the British Museum. Annals and Magazine of Natural History (Series 7) v. 12 (núm. 72): 621-630