Les seues principals amenaces són la desforestació, les pràctiques agrícoles gens sostenibles i la construcció de preses que redueixen o interrompen el cabal dels rius.[14]
↑McClelland J., 1838. Observations on six new species of Cyprinidae, with an outline of a new classification of the family. J. Asiatic Soc. Bengal v. 7 (for Nov. 1838). 941-948.
↑ 2,02,1 Mai, D. Y., 1978. Identification of the fresh-water fishes of North Viet Nam. Ha Noi, Scientific & Technology, Publisher. Freshwater Fish Vietnam: 1-340, Pls. 1-48.
↑Chu, X.-L. i Chen, Y.-R., 1990. The Fishes of Yunnan, China. Part II. Science press, Beijing.
↑Kottelat, M., 2001. Freshwater fishes of northern Vietnam. A preliminary check-list of the fishes known or expected to occur in northern Vietnam with comments on systematics and nomenclature. Environment and Social Development Unit, East Asia and Pacific Region. El Banc Mundial. Freshwater Fish. El Vietnam (BOOK), juny: i-iii + 1-123 + 1-18.
↑Kottelat, M., 1989. Zoogeography of the fishes from Indochinese inland waters with an annotated check-list. Bull. Zoöl. Mus. Univ. Amsterdam 12(1): 1-55.
Banarescu, P. i T. T. Nalbant 1995: A generical classification of Nemacheilinae with description of two new genera (Teleostei: Cypriniformes: Cobitidae). Travaux du Museum d'Histoire Naturelle "Grigore Antipa" v. 35: 429-495.
Jayaram, K. C. 1981: The freshwater fishes of India, el Pakistan, Bangladesh, Burma and Sri Lanka--a handbook: i-xxii + 1-475, Pls. 1-13. Zoological Survey of India.
Moyle, P.; Cech, J. Fishes: An Introduction to Ichthyology (en anglès). 4a edició. Prentice Hall, 2000. ISBN 9780130112828.
Nelson, J. Fishes of the World (en anglès). 3a edició. John Wiley and Sons, 1994. ISBN 978-0471547136.