Boulenger, 1894 : List of Reptiles and Batrachians collected by Dr. J. Bohls near Asuncion, Paraguay. Annals and Magazine of Natural History, sér. 6, vol. 13, no 76, p. 342-348 (texte intégral).
↑de Sá, Grant, Camargo, Heyer, Ponssa & Stanley, 2014 : Systematics of the Neotropical genus Leptodactylus Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the relevance of non-molecular evidence, and species accounts. South American Journal of Herpetology, vol. 9, no 1, p. 1–128 (texte intégral)