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Endomychidae

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Endomychidae
Temporal range: Cenomanian–Recent
Brachytrycherus humeralis
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Arthropoda
Clade: Pancrustacea
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Infraorder: Cucujiformia
Superfamily: Coccinelloidea
Family: Endomychidae
Leach, 1815
Type genus
Endomychus
Synonyms

Merophysiidae Seidlitz, 1872

Endomychidae, also commonly known as handsome fungus beetles, is a family of coccinelloid beetles with representatives found in all biogeographic realms. As the common name suggests, members of this family feed on fungi.

Description

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They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment.

Taxonomy

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The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea.

Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.[2]

This family contains around 120 genera and 1300 species. A list of subfamilies can be seen below:

The subfamilies that are included:

Genera

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Genera traditionally placed in family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):[verification needed]

Data sources: i = ITIS,[3] c = Catalogue of Life,[4] g = GBIF,[5] b = Bugguide.net[6]

Mimolithophilus and Monocoryna are now placed in Coccinellidae.

Fossil genera

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References

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  1. ^ Tomaszewska, K.Wioletta (2000). "Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)" (PDF). Annales Zoologici. 50 (4): 449–558.
  2. ^ Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B; Whiting, Michael F; McHugh, Joseph V (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)" (PDF). Systematic Entomology. 40 (4): 745. Bibcode:2015SysEn..40..745R. doi:10.1111/syen.12138. S2CID 55206626.
  3. ^ "Endomychidae Report". Integrated Taxonomic Information System. Retrieved 2018-05-07.
  4. ^ "Browse Endomychidae". Catalogue of Life. Archived from the original on 2021-06-18. Retrieved 2018-05-07.
  5. ^ "Endomychidae". GBIF. Retrieved 2018-05-07.
  6. ^ "Endomychidae Family Information". BugGuide.net. Retrieved 2018-05-07.
  7. ^ a b c d e Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018). "The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber". Cretaceous Research. 91: 287–298. Bibcode:2018CrRes..91..287T. doi:10.1016/j.cretres.2018.07.001. S2CID 135401188.
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