Copelatus

Copelatus
Thời điểm hóa thạch: Eocene[1] - nay
Phân loại khoa học
Giới (regnum)	Animalia
Ngành (phylum)	Arthropoda
Lớp (class)	Insecta
Bộ (ordo)	Coleoptera
Họ (familia)	Dytiscidae
Phân họ (subfamilia)	Copelatinae
Chi (genus)	Copelatus Erichson, 1832

Copelatus là một chi bọ cánh cứng trong họ Dytiscidae. Có khoảng 470 loài đã được miêu tả, được tìm thấy trên toàn cầu, nhưng chúng đa dạng nhất ở vùng nhiệt đới của Nam Mỹ, Châu Phi và Đông Nam Á.

Loài

Copelatus được chia thành một số phân chi (Colepatus, Papuadytes etc.), một vài phân chia đôi khi được xem là các chi riêng biệt. Nó có thể là cận ngành với các chi trong phân họ Copelatinae là Lacconectus và Aglymbus^[2]. Các loài bao gồm:

Copelatus abonnenci Guignot, 1939 Copelatus acamas Guignot, 1955 Copelatus advena Sharp, 1882 Copelatus aemulus Bilardo & Rocchi, 1995 Copelatus aequatorius Régimbart, 1899 Copelatus aethiopicus Régimbart, 1906 Copelatus agrias Guignot, 1954 Copelatus aldabricus J.Balfour-Browne, 1950 Copelatus alternatus Sharp, 1882 Copelatus amaroides Guignot, 1952 Copelatus amatolensis Omer-Cooper, 1965 Copelatus amazonicus Régimbart, 1889 Copelatus ambiguus Bertrand & Legros, 1975 Copelatus anastomosans Guignot, 1952 Copelatus andobonicus Guignot, 1960 Copelatus andreinii Régimbart, 1905 Copelatus andrewesi J.Balfour-Browne, 1939 Copelatus angolensis Peschet, 1924 Copelatus angustus Gschwendtner, 1932 Copelatus annobomensis J.Balfour-Browne, 1939 Copelatus anthracinus Régimbart, 1895 Copelatus aphroditae Balke, 2003 Copelatus apicalis Fairmaire, 1898 Copelatus apuzzoi Bilardo & Rocchi, 1999 Copelatus aruensis J.Balfour-Browne, 1939 Copelatus assamensis Vazirani, 1970 Copelatus assimilis Régimbart, 1895 Copelatus ateles Guignot, 1955 Copelatus atrosulcatus Régimbart, 1906 Copelatus bacchusi Wewalka, 1981 Copelatus bacillifer Guignot, 1955 Copelatus baculiformis Guignot, 1955 Copelatus badeni Sharp, 1882 Copelatus bakewelli J.Balfour-Browne, 1939 Copelatus bangalorensis Vazirani, 1970 Copelatus baoulicus Bilardo & Pederzani, 1978 Copelatus barbouri Young, 1942 Copelatus basifasciatus Régimbart, 1895 Copelatus basilewskyi Bilardo & Pederzani, 1979 Copelatus bechynei Guignot, 1953 Copelatus befasicus Guignot, 1956 Copelatus bengalensis Guignot, 1955 Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996 Copelatus bibulus Guignot, 1948 Copelatus biformis Sharp, 1882 Copelatus bilunatus Guignot, 1955 Copelatus binaghii Bilardo & Pederzani, 1978 Copelatus biolleyi Guignot, 1952 Copelatus biroi Guignot, 1956 Copelatus biseriatus J.Balfour-Browne, 1939 Copelatus biswasi Mukherjee & Sengupta, 1986 Copelatus blancasi Guignot, 1958 Copelatus blatchleyi Young, 1953 Copelatus bolivianus Guignot, 1957 Copelatus bombycinus Guignot, 1952 Copelatus bonvouloiri Sharp, 1882 Copelatus bottegoi Régimbart, 1895 Copelatus boukali Hendrich & Balke, 1998 Copelatus brancuccii Rocchi, 1979 Copelatus brasiliensis Zimmermann, 1921 Copelatus brevistrigatus Guignot, 1959 Copelatus brivioi Rocchi, 1976 Copelatus brullei Aubé, 1838 Copelatus brunneus J.Balfour-Browne, 1939 Copelatus buqueti Aubé, 1838 Copelatus burgeoni Gschwendtner, 1930 Copelatus caelatipennis Aubé, 1838 Copelatus caelatus Guignot, 1952 Copelatus caffer J.Balfour-Browne, 1939 Copelatus camerunensis Guignot, 1941 Copelatus capensis Sharp, 1882 Copelatus carayoni Legros, 1949 Copelatus carinatus Sharp, 1882 Copelatus celinoides Guignot, 1952 Copelatus ceylonicus Vazirani, 1969 Copelatus cheesmanae J.Balfour-Browne, 1939 Copelatus chevrolati Aubé, 1838 Copelatus chibcha Guignot, 1952 Copelatus chipiriricus Guignot, 1957 Copelatus chloroticus Régimbart, 1899 Copelatus cinnamomeus Régimbart, 1895 Copelatus clarki Sharp, 1882 Copelatus collarti Gschwendtner, 1932 Copelatus compertus Guignot, 1956 Copelatus concii Bilardo, 1982 Copelatus concolor Sharp, 1882 Copelatus concolorans J.Balfour-Browne, 1939 Copelatus confinis Bilardo & Rocchi, 1999 Copelatus congo Gschwendtner, 1938 Copelatus consimilis Bilardo & Rocchi, 2002 Copelatus consors Sharp, 1882 Copelatus cooperae Bilardo & Pederzani, 1972 Copelatus cordylinoides J.Balfour-Browne, 1938 Copelatus coxalis Sharp, 1882 Copelatus crassus Régimbart, 1895 Copelatus cryptarchoides Régimbart, 1899 Copelatus cubaensis Schaeffer, 1908 Copelatus curtistriatus Bilardo & Rocchi, 1995 Copelatus daemeli Sharp, 1882 Copelatus darlingtoni Young, 1942 Copelatus debilis Sharp, 1882 Copelatus decellei Bilardo, 1982 Copelatus decemsulcatus Régimbart, 1895 Copelatus deceptor Bilardo & Rocchi, 1995 Copelatus depressus Sharp, 1882 Copelatus descarpentriesi Bertrand & Legros, 1975 Copelatus diffisus Guignot, 1939 Copelatus dimorphus Sharp, 1882 Copelatus distinctus Aubé, 1838 Copelatus distinguendus Régimbart, 1903 Copelatus divisus Watts, 1978 Copelatus dolosus Guignot, 1956 Copelatus doriae Sharp, 1882 Copelatus duodecimstriatus Aubé, 1838 Copelatus duponti Aubé, 1838 Copelatus edax Guignot, 1955 Copelatus efoutensis Bilardo & Rocchi, 1995 Copelatus ejactus Omer-Cooper, 1965 Copelatus ellai Bilardo & Rocchi, 1995 Copelatus elutus Guignot, 1958 Copelatus enganensis Guignot, 1940 Copelatus epactus Guignot, 1948 Copelatus erichsonii Guérin-Méneville, 1849 Copelatus esteriensis Bilardo & Pederzani, 1978 Copelatus eucritus Guignot, 1952 Copelatus evanidus Bilardo & Rocchi, 1995 Copelatus exaratus Sharp, 1882 Copelatus externus Kirsch, 1873 Copelatus fasciatus Bilardo & Rocchi, 1995 Copelatus fastidiosus Guignot, 1959 Copelatus feae Régimbart, 1888 Copelatus fernandensis Guignot, 1952 Copelatus ferruginicollis Régimbart, 1895 Copelatus ferus Guignot, 1953 Copelatus festae Régimbart, 1899 Copelatus fidschiensis Zimmermann, 1928 Copelatus fijiensis Guignot, 1955 Copelatus filiformis Sharp, 1882 Copelatus flavicans Guignot, 1952 Copelatus flavidus Régimbart, 1895 Copelatus fluviaticus Guignot, 1957

Copelatus fontanus J.Balfour-Browne, 1950 Copelatus fossilis Riha, 1974 Copelatus fractistriatus Bilardo & Rocchi, 1995 Copelatus freudei Guignot, 1955 Copelatus fryi J.Balfour-Browne, 1939 Copelatus fulviceps J.Balfour-Browne, 1938 Copelatus fuscipennis Sharp, 1882 Copelatus fuscomaculatus Guignot, 1952 Copelatus gabonicus Bilardo & Pederzani, 1978 Copelatus galapagoensis G.R.Waterhouse, 1845 Copelatus garambanus Guignot, 1955 Copelatus gardineri Scott, 1912 Copelatus geayi Régimbart, 1904 Copelatus geniculatus Sharp, 1882 Copelatus gentilis Sharp, 1882 Copelatus gestroi Régimbart, 1892 Copelatus gibsoni Vazirani, 1974 Copelatus glyphicus (Say, 1823) Copelatus griffinii Régimbart, 1899 Copelatus gschwendtneri Guignot, 1939 Copelatus guadelupensis Legros, 1948 Copelatus guerini Aubé, 1838 Copelatus guineensis Guignot, 1955 Copelatus hararensis Guignot, 1952 Copelatus hardenbergi J.Balfour-Browne, 1950 Copelatus hebeter Guignot, 1953 Copelatus heterogynus Régimbart, 1899 Copelatus hydroporoides (Murray, 1859) Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000 Copelatus iguelaensis Bilardo & Rocchi, 2002 Copelatus ilybioides Régimbart, 1895 Copelatus imasakai Matsui & Kitayama, 2000 Copelatus imitator Bilardo & Rocchi, 2002 Copelatus inaequalis Sharp, 1882 Copelatus incognitus Sharp, 1882 Copelatus indicus Sharp, 1882 Copelatus inopinatus Bilardo & Rocchi, 1995 Copelatus inornatus Sharp, 1882 Copelatus insidiosus Bilardo & Rocchi, 1995 Copelatus insolitus Chevrolat, 1863 Copelatus instabilis Régimbart, 1897 Copelatus instriatus Zimmermann, 1921 Copelatus insuetus Guignot, 1941 Copelatus insulanus Guignot, 1939 Copelatus integer Sharp, 1882 Copelatus internus Régimbart, 1904 Copelatus interstriatus J.Balfour-Browne, 1939 Copelatus inuber Guignot, 1952 Copelatus ipiformis Régimbart, 1895 Copelatus irinus Régimbart, 1899 Copelatus irregularis W.J. Macleay, 1871 Copelatus ischius Guignot, 1956 Copelatus jactator Guignot, 1955 Copelatus jamaicensis Young, 1942 Copelatus japonicus Sharp, 1884 Copelatus jarrigei Legros, 1954 Copelatus javanus Régimbart, 1883 Copelatus jocosus Guignot, 1955 Copelatus johannis Guignot, 1954 Copelatus kalaharii Gschwendtner, 1935 Copelatus kammuriensis Tamu & Tsukamoto, 1955 Copelatus karnatakus Holmen & Vazirani, 1990 Copelatus kaszabi Guignot, 1956 Copelatus kilimandjarensis Bilardo & Pederzani, 1972 Copelatus kindianus Guignot, 1954 Copelatus kongouensis Bilardo & Rocchi, 1999 Copelatus koreanus Mori, 1932 Copelatus laccophilinus Sharp, 1882 Copelatus laeticulus Sharp, 1882 Copelatus lamottei Legros, 1954 Copelatus lanzai Bilardo & Rocchi, 1995 Copelatus laraensis Bilardo & Rocchi, 1995 Copelatus lasckonyi Bilardo & Rocchi, 1995 Copelatus latens Guignot, 1952 Copelatus laticollis Régimbart, 1899 Copelatus latifasciatus Bilardo & Rocchi, 1999 Copelatus latipes Sharp, 1882 Copelatus latus J.Balfour-Browne, 1950 Copelatus leonardii Bilardo & Rocchi, 1999 Copelatus leonensis Legros, 1954 Copelatus lepersonneae Gschwendtner, 1943 Copelatus lignosus Guignot, 1955 Copelatus lineatipennis Guignot, 1955 Copelatus lineatus (Guérin-Méneville, 1838) Copelatus longicornis Sharp, 1882 Copelatus lootensi Guignot, 1955 Copelatus louayensis Bilardo & Rocchi, 2004 Copelatus luctuosus Guignot, 1939 Copelatus luridescens Régimbart, 1889 Copelatus luteocinctus Guignot, 1955 Copelatus luteomaculatus Guignot, 1956 Copelatus luzonicus Guignot, 1952 Copelatus macellus Guignot, 1950 Copelatus madoni Guignot, 1952 Copelatus mahajanga Pederzani & Hájek, 2005 Copelatus mahleri Holmen & Vazirani, 1990 Copelatus makokouensis Bilardo & Rocchi, 1995 Copelatus malaisei Guignot, 1954 Copelatus mancus Sharp, 1887 Copelatus marginatus Sharp, 1882 Copelatus masculinus Régimbart, 1899 Copelatus massaicus Guignot, 1941 Copelatus mathani Guignot, 1952 Copelatus mbokoensis Bilardo & Rocchi, 2006 Copelatus melanogrammus Régimbart, 1883 Copelatus mimetes Guignot, 1957 Copelatus minimus Bilardo & Rocchi, 1995 Copelatus minor Bilardo & Pederzani, 1978 Copelatus minutissimus J.Balfour-Browne, 1939 Copelatus mocquerysi Régimbart, 1895 Copelatus mohelicus Guignot, 1959 Copelatus monticola Guignot, 1951 Copelatus montivagus Young, 1942 Copelatus mulangensis Bameul, 2003 Copelatus mundus Sharp, 1882 Copelatus mutabilis Bilardo & Rocchi, 1999 Copelatus mvoungensis Bilardo & Rocchi, 2004 Copelatus mysorensis Vazirani, 1970 Copelatus nakamurai Guéorguiev, 1970 Copelatus nangaensis Guignot, 1952 Copelatus nauclerus Guignot, 1939 Copelatus neavei J.Balfour-Browne, 1950 Copelatus neelumae Vazirani, 1973 Copelatus neoguineensis Zimmermann, 1919 Copelatus nigricans Sharp, 1882 Copelatus nigrolineatus Sharp, 1882 Copelatus nigropennis Zimmermann, 1927 Copelatus nigrostriatus Régimbart, 1895 Copelatus nilssoni Wewalka, 1982 Copelatus nimbaensis Legros, 1958 Copelatus nitens Bilardo & Rocchi, 1999 Copelatus nitidus Sharp, 1882 Copelatus nodieri Régimbart, 1895 Copelatus normalis Erichson, 1847 Copelatus notius Omer-Cooper, 1965 Copelatus nzei Bilardo & Rocchi, 1999 Copelatus oblitus Sharp, 1882 Copelatus obscurus Sharp, 1882 Copelatus occultus Bilardo & Rocchi, 1995 Copelatus ogasawarensis Kamiya, 1932 Copelatus onorei Pederzani & Rocchi, 1982

Copelatus ornatipennis Zimmermann, 1926 Copelatus owas Régimbart, 1895 Copelatus pachys Guignot, 1955 Copelatus pallidus Régimbart, 1895 Copelatus paludorum Guignot, 1952 Copelatus pandanorum Scott, 1912 Copelatus pantosi Guignot, 1958 Copelatus papuensis J.Balfour-Browne, 1939 Copelatus parabaptus Guignot, 1955 Copelatus parallelipipedus Régimbart, 1895 Copelatus parallelus Zimmermann, 1920 Copelatus pardii Rocchi, 1990 Copelatus parisii Guignot, 1959 Copelatus parumstriatus Gschwendtner, 1934 Copelatus paryphes Guignot, 1955 Copelatus pederzanii Bilardo & Rocchi, 1995 Copelatus pereirai Guignot, 1955 Copelatus peridinus Guignot, 1955 Copelatus pinnifer Guignot, 1952 Copelatus piriensis Omer-Cooper, 1965 Copelatus platynotus Régimbart, 1895 Copelatus polystrigus Sharp, 1882 Copelatus ponomarenkoi Riha, 1974 Copelatus portior Guignot, 1956 Copelatus posticatus (Fabricius, 1801) Copelatus poungai Bilardo & Rocchi, 2006 Copelatus predaveterus K.B.Miller, 2003 Copelatus prolixus Sharp, 1882 Copelatus prolongatus Sharp, 1882 Copelatus propino Guignot, 1960 Copelatus propinquus Régimbart, 1895 Copelatus proximus Sharp, 1882 Copelatus pulchellus (Klug, 1834) Copelatus pulicarius Régimbart, 1895 Copelatus pumilus Régimbart, 1895 Copelatus punctatus Bilardo & Rocchi, 1995 Copelatus punctulatus Aubé, 1838 Copelatus quadrisignatus Régimbart, 1877 Copelatus racenisi Guignot, 1951 Copelatus ragazzii Régimbart, 1887 Copelatus regimbarti Branden, 1885 Copelatus restrictus Sharp, 1882 Copelatus rimosus Guignot, 1952 Copelatus rivalis Guignot, 1952 Copelatus rocchii Bilardo, 1982 Copelatus romani Zimmermann, 1924 Copelatus royi Legros, 1958 Copelatus rubiginosus Guignot, 1954 Copelatus ruficapillus Régimbart, 1895 Copelatus ruteri Legros, 1954 Copelatus saegeri Guignot, 1955 Copelatus sahlbergi J.Balfour-Browne, 1939 Copelatus sallaei Sharp, 1882 Copelatus sanfilippoi Bilardo & Pederzani, 1972 Copelatus scalptus Guignot, 1954 Copelatus scaphites Guignot, 1955 Copelatus schaefferi Young, 1942 Copelatus schereri Wewalka, 1981 Copelatus schuhi Hendrich & Balke, 1998 Copelatus scutatus Guignot, 1952 Copelatus scytalotus Guignot, 1956 Copelatus senegalensis Lagar & Hernando, 1991 Copelatus sexstriatus Sharp, 1882 Copelatus sharpi Branden, 1885 Copelatus silvestrii Régimbart, 1903 Copelatus simoni Régimbart, 1889 Copelatus singularis Bilardo & Rocchi, 1995 Copelatus sociennus J.Balfour-Browne, 1952 Copelatus solitarius Sharp, 1882 Copelatus sordidipennis Régimbart, 1895 Copelatus spangleri Vazirani, 1974 Copelatus speciosus Régimbart, 1892 Copelatus spoliatus Guignot, 1955 Copelatus stavropolitanus Riha, 1974 Copelatus stillicidicola Bilardo & Rocchi, 1995 Copelatus striatellus Boheman, 1848 Copelatus striaticollis Lucas, 1857 Copelatus striatopterus Aubé, 1838 Copelatus striatulus Aubé, 1838 Copelatus strigipennis (Laporte, 1835) Copelatus strigosulus (Fairmaire, 1878) Copelatus strinatii Guignot, 1958 Copelatus striolatus Peschet, 1917 Copelatus stygis Guignot, 1958 Copelatus subdeficiens Régimbart, 1902 Copelatus subsimilis Guignot, 1958 Copelatus substriatus Kirsch, 1873 Copelatus subterraneus Guéorguiev, 1978 Copelatus sudrei Bameul, 2003 Copelatus sulcatus Sharp, 1882 Copelatus sulcipennis (Laporte, 1835) Copelatus sumbawensis Régimbart, 1899 Copelatus suppar Guignot, 1956 Copelatus supplementaris Régimbart, 1895 Copelatus sylvaticus Guignot, 1952 Copelatus takakurai Satô, 1985 Copelatus tanaus Guignot, 1953 Copelatus taprobanicus Wewalka & Vazirani, 1985 Copelatus tenebrosus Régimbart, 1880 Copelatus teranishii Kamiya, 1938 Copelatus terminalis Sharp, 1882 Copelatus ternatensis Régimbart, 1899 Copelatus thiriouxi Peschet, 1917 Copelatus thrasys Guignot, 1952 Copelatus tibialis Sharp, 1882 Copelatus tinctor Guignot, 1954 Copelatus togoensis Régimbart, 1895 Copelatus tomokunii Satô, 1985 Copelatus tostus J.Balfour-Browne, 1950 Copelatus trifilis Guignot, 1958 Copelatus triglyphus Guignot, 1955 Copelatus trilobatus Régimbart, 1895 Copelatus tschaga Bilardo & Pederzani, 1972 Copelatus tucuchiensis J.Balfour-Browne, 1939 Copelatus tulagicus Guignot, 1942 Copelatus uludanuensis Hendrich & Balke, 1995 Copelatus undecimstriatus Aubé, 1838 Copelatus unguicularis Régimbart, 1903 Copelatus usagarensis Zimmermann, 1926 Copelatus vagatus Guignot, 1952 Copelatus vagestriatus Zimmermann, 1919 Copelatus vagus Bilardo & Rocchi, 1995 Copelatus validus Sharp, 1882 Copelatus vanninii Bilardo & Rocchi, 1999 Copelatus variegatus Régimbart, 1895 Copelatus venustus Bilardo & Rocchi, 1995 Copelatus vigintistriatus Fairmaire, 1869 Copelatus vigintisulcatus Régimbart, 1895 Copelatus villiersi Guignot, 1950 Copelatus virungaensis Bilardo, 1982 Copelatus vitraci Legros, 1948 Copelatus vivax Guignot, 1953 Copelatus wallacei J.Balfour-Browne, 1939 Copelatus waltoni J.Balfour-Browne, 1950 Copelatus wewalkai Holmen & Vazirani, 1990 Copelatus weymarni J.Balfour-Browne, 1947 Copelatus xanthocephalus Régimbart, 1899 Copelatus yacumensis Guignot, 1957 Copelatus zadiensis Bilardo & Rocchi, 1995 Copelatus zimmermanni Gschwendtner, 1934

Hình ảnh

• 

Chú thích

1. Miller,K.B, Blake,M. The unusual occurrence of aquatic beetles in amber, copelatus aphroditae balke, n. sp. and C. predaveterus miller, n. sp., (Coleoptera: Dytiscidae: Copelatinae). Proc. of the Entomological Society of Washington, 2003, vol. 105, no4, pp. 809-815
2. Balke,M et al., MtDNA phylogeny and biogeography of Copelatinae, a highly diverse group of tropical diving beetles (Dytiscidae). Molecular Phylogenetics and Evolution Vol. 32, Issue 3, 2004

Tham khảo